CARABUS EUROPE + TURKEY


europe subgenus tween = + palalearct, nearct






CARABUS Linné, 1758;
species: 7



1. arvensis Herbst, 1784; Europe, Siberia, Far East, Japan


ssp. arvensis Herbst, 1784; emend. for arcensis Herbst N,C-Europe:Slovenia to Scandinavia; W-Eur.Russia; "Pommern"(=NW-Poland)
~arcensis Paykull, 1790; ios of arvensis Herbst
=pomeranus Gmelin, 1788 "Pommern"(=currently NW-Poland)
~aereus Dejean, 1826, nec Dejean, 1826 "Alps of Steiermark"
=seileri Heer, 1837 Switzerland:Schaffhausen
~cupreoaeneus Letzner, 1850; un
~marginatus Letzner, 1850; un
~niger Letzner, 1850; un
~nigrescens Letzner, 1850; un
~purpurescens Letzner, 1850; un
~versicolor Letzner, 1850; un
~viridiaeneus Letzner, 1850; un
~viridis Letzner, 1850; un
~ziegleri Géhin, 1876, nec Duftschmid, 1812; nom.pro aereus Dejean "Alps of Steiermark"
~cuprescens Dalla Torre, 1877; un
~fuscescens Dalla Torre, 1877; un
~virescens Dalla Torre, 1877; un
~nigrescens Westhoff, 1881; un
~nigrinus Westhoff, 1881, nec Motschulsky, 1865
~aeratus Géhin, 1885, nec Quensel, 1806; nom.pro ziegleri Géhin "Alps of Steiermark"
~ruficrus Géhin, 1885; un
~nigrinopomeranus Rade, 1895; un
=austriae Sokolar, 1907 ?Wienerwald
~detritus Lapouge, 1908, nec Drapiez, 1821 N-Germany:?
~simplex Lapouge, 1908; un "S-Germany & Austria"
=noricus Sokolar, 1910 Austria:Alps:Wechsel
=decipiens Lengerken, 1911 "Bayern"
=germaniae Lengerken, 1911 Germany:?
=borussicus Csiki, 1927; nom.pro detritus Lapouge N-Germany:?
~baldiai Mandl, 1963; un Austria:Vorarlberg:Hochfreschen, 1900m
~haideri Mandl, 1963; un Kärnten:Schleinitz, Lienzer Hütte env.

ssp. conciliator Fischer, 1822 E-Siberia(Nerchinsk), Baykal Lake; N-Mongolia; Altays, Sayans
=dahuricus Gebler, 1827 ?"Dauria"
=kirbyi Fischer, 1827 Lena River, Taymyr Peninsula
=vinculatus Gebler, 1830 Barnaul
=kamtchaticus Motschulsky, 1846 Kamchatka(LT)
~borealis Poppius, 1906, nec Paykull, 1790 lower Lena River:"Shigansk"
~jelanensis Lapouge, 1908; un "Jelan"(=Baykal Lake env.)
=poppiusi Csiki, 1927; nom.pro borealis Poppius lower Lena River:"Shigansk"
=jurgitae Deuve, 1993 Altays Mts.:Aktash

ssp. faldermanni Dejean, 1829 Far East, Korea; Shanxi, Shaanxi & NE-China; "Dauria"
=femoralis Motschulsky, 1865; nec ? "Amur"(LT)
=amurensis Géhin, 1885; nom.pro femoralis Motschulsky "Amur"(LT)
=bellus Kraatz, 1886
=concinnus Kraatz, 1886 ?"Amur"
=nigerrimus Kraatz, 1886
=viridicoeruleus Kraatz, 1886
=viridiniger Kraatz, 1886 ?"Amur"
=provostii Fairmaire, 1888 "Mts.nr.Peking"
=praesensum Cavazzuti, 1999 C-Shaanxi:Tongchuan:Shimen Shan, S-slope, 1300-1600m; 1m/1f

ssp. hokkaidoensis Lapouge, 1924 Japan:Hokkaido("Teshio"); Kurils; Sakhalin
=sachalinensis Lapouge, 1906 "Sakhalin"

ssp. baschkiricus Breuning, 1936 CE-Russia(Kazan’); C-Urals
~laticollis Motschulsky, 1850, nec Sturm, 1826 Baschkiria:Mt.Ilmen(=?Ilmen’ ca.20km NE Zlatoust)

ssp. sylvaticus Dejean, 1826 W-Europe:France(Seine-Marne:Ft.d’Eu), W-Switz., Belgium, Holland, W-Germany, Brit.Isles
=schrickelli Dejean, 1826 LNF
=alpicola Heer, 1837 ?Switzerland:Mt.Pilatus
=anglicus Motschulsky, 1865 Windsore nr.London
~irregularis Beuthin, 1896, nec Fabricius, 1792 Haute-Loire
=raetzeri Born, 1898 Switzerland:Neuenburger Jura
~costalis Lapouge, 1908; un France:Cévennes:Gerbier des Jonc
=liebmanni Lengerken, 1911 W-France:Vosges
=spencei Lengerken, 1911 "Pyrenees"(error)
=sugintensis Brunier, 1922 W-France:Haut-Rhin:Sundgau(=area betw.Basel/Mulhouse)
=subsylvaticus Perraudin, 1965 ?"Stoccarda"
=jarrigei Colas, 1971 Cote d’Or:Chatillon; TNF
=thebaudi Colas, 1971 Corréze:Les Monédiéres; TNF
~bicolor (Keith, 1989); un
=couchyi Freney, 2004 ?

ssp. carpathus Born, 1902; ?non priority Carpathians(C-Romania:Bucegi Mt.R., Mt.Negoi); E-Europe, S-Eur.Russia
=?eremita Fischer, 1823; ?priority "Kaukasus"(error)
~romanus Marcu, 1937; un Romania:?
=csikensis Csiki, 1942 CE-Romania:Haghimas Mts.nr.Balan
=pseudonoricus Kenyery, 1983 Slovenia:"Bachergebirge:N-slope:1km S Podvelka, 350m"(=ca.betw.Celje/Zagreb)

ssp. venetianus Bernau, 1914 NE-Italy:Dolomiti, Veneto(Bosco di Cansiglio); Slovenia



2.deyrollei Gory, 1839 NW-Spain(Galicia:Sierra de Perache); N-Portugal
~luctuosus Géhin, 1885, nec Villers, 1789 N-Portugal:Braga
~chloromorphus Branes, 1985; un
~cyaneomorphus Branes, 1985; un
~goryi Branes, 1985; un
~miniresplendens Branes, 1985; un
~neotristis Branes, 1985; un
~pecoudi Branes, 1985; un
~splendidulus Branes, 1985; un



3.granulatus Linné, 1758 Europe, Brit.Isles, Balcans, Turkey, Siberia, Far East, Japan; introduced: Canada, USA

ssp. granulatus Linné, 1758 C,W,N-Europe, Brit.Isles, Balcans; S-Russia; introd.:SW-Canada:Brit.Columbia; ?LNF
=chalybaeus Voet, 1776
=campestris Adams, 1817 Volga River Bank
=confluens Fischer, 1827 Volga River
=parallelus Faldermann, 1835 Volga River
~niger Letzner, 1850; un
~nigrescens Letzner, 1850; un
~rufofemoratus Letzner, 1850; un
~virescens Letzner, 1850; un
~rubripes Géhin, 1876, nec Duftschmid, 1812
~cupreoaeneus Dalla Torre, 1877; un
~nigroaeneus Dalla Torre, 1877; un
~viridiaeneus Dalla Torre, 1877; un
=forticostis Kraatz, 1878
=haematomerus Kraatz, 1878
=parvicollis Kraatz, 1878 ?"Steiermark"
~fulvipes Géhin, 1885, nec Fabricius, 1792
=wimmeli Schultz, 1900 Hamburg
~hudsonicus Lapouge, 1924, nec Motschulsky, 1865 ?Hudson Bay
=granulatulus Mandl, 1955
~crimeensis Breuning, 1933; un Krym Peninsula:?Jaila Mts.
=hibernicus Lindroth, 1956 Ireland:Kerry Co.:Killarney; Hebrides; introduced:Quebec, N.Brunswick, N.Scotia
~atrocyanescens Lamy, 1973; un

ssp. duarius Fischer, 1844; nom.pro elongatus Fischer Siberia, Altays, Sayans, Kazakhstan, Kyrgyzstan, N-Mongolia; "Dzhungaria"
~elongatus Fischer, 1842, nec Duftschmid, 1812 "Dzhungaria"
=songaricus Motschulsky, 1850 "Des.Kirg.or."(LT)
=dauricus Solsky, 1875 Irkutsk & Krasnoyarsk
~solskyanus Géhin, 1885; un
=cupriculus Reitter, 1896 Siberia
=lenensis Poppius, 1906 lower Lena River
~luctuosus Lapouge, 1924, nec Villers, 1789 "Tunkun-Sayan"(=Siberia/Mongolia:Tunkunskiye Goltsy Mts.)
=luctisonius Csiki, 1927; nom.pro luctuosus Lapouge "Tunkun-Sayan"(=Siberia/Mongolia:Tunkunskiye Goltsy Mts.)


ssp. interstitialis Duftschmid, 1812 S-Austria(Sudkärnten); Italy, Corsica, S-Balcans; Turkey:Konya, Denizli, Mugla, Samsun
=palustris Dejean, 1826 Roma env.
=aetolicus Schaum, 1857 C-Greece:"Missolunghi"(=Mesolóngi)
=debilicostis Kraatz, 1878 ?Piemonte
=miriditus Apfelbeck, 1901 N-Albania:"Velipoja nr.Alessio"(=Lezhë)
=paludicola Müller, 1903 Hercegovina:Narenta River nr.Metkovich
=corsicus born, 1906 Corsica:Folelli
=calabricus Spettoli & Vigna Taglianti, 2001 S-Italy:Calabria:Parco regoinale delle Serre:Colle del Monaco

ssp. corticalis Motschulsky, 1846, nec Fabricius, 1801; nbm S-Dagestan, NE-Georgia, N-Azerbaydzhan; "Georgia";"Kachetia:Matany"(LT)
~ponticus Lapouge, 1924, nec Deyrolle, 1869 "Amasya"(?by error, LT)
=ponti Csiki, 1927; nom.pro ponticus Lapouge "Amasya"(?by error, LT)
=daghestanicus Lapouge, 1924 Dagestan

ssp. leander Kraatz, 1878 C-Caucasus:N-macroslope:Teberda to W-Dagestan; ?LNF

ssp. yezoensis Bates, 1883 Japan:Hokkaido("Sapporo & Junsai Lake nr.Hakodate"); Sakhalin
~sachalinensis Matsumura, 1911, nec Lapouge, 1906 Sakhalin
~praedator Lapouge, 1924; un "Yesso"(=Hokkaido)
=karafutensis Csiki, 1927; nom.pro sachalinensis Matsumura Sakhalin

ssp. telluris Bates, 1883 Japan:N,C-Honshu(Tonosawa); NE-China, Shanxi, Far East, Korea
=pekinensis Fairmaire, 1887 "Peking"
=expansus Lapouge, 1905 Selenga River
=ussuriensis Born, 1914 Sikhote-Alin Mt.R.
~gobiensis Lapouge, 1924; un "Inn-Shan & Kalgan"(=Hebei:Xianhua)



4.menetriesi Hummel, 1827 C-Europe, NE-Europe, N-Russia, Finland, Byelorussia, Latvia

ssp. menetriesi Hummel, 1827 N-Europe, Finland, N-Russia(Novgorod env.), Byelorussia, Latvia; N-Slovakia:Oravská Lesná


ssp. pacholei Sokolar, 1911 Austria:Oberöst.(Tanner Moor nr.Königswiesen); S,W,N-Bohemia: Šumava Mts. to Ústí n.Lab.
~bohemicus Tanzer, 1934; un S-Bohemia:"Mt.Schreiner foot:Schneider- & Oberschlag"(=Mt.Bobík:Krejèovice, Milešice)
=knabli Mandl, 1951 E-Austria:"Tirol:Neumühl(=Breitenwang) nr.Reutte"(=ca.60km WNW Innsbruck); 1f
=pseudogranulatus Nüssler, 1965 Germany:Sachsen:Gayer:Hermannsdorfer Wiese
=witzgalli Reiser in Freude, 1976; ?nud



5.sculpturatus Ménétriés, 1832 SE-Azerbaydzhan:Lenkoran’, Talysh Mts.; N-Iran:Gilan, Elborz Mts., Gorgan env.
=lenkoranus Reitter, 1896 ?"Lenkoran"
=gilanicus Mandl, 1967 Gilan:?betw.Asalam/Heroabad
=hyrcanus Heinz, 1977 Mazanderan:30km SE Beshar



6.stscheglowi Mannerheim, 1827 E-Ukraine, E-Eur.Russia; C,S-Urals(Zlatoust betw.Ufa/Chelyabinsk)
=zakharschewskii Motschulsky, 1845 Ukraine:"Charcov:Tschuguev"(=Kharkov:Chuguevka); "Rus.mer.:Svatova Luchka"(LT)



7.vanvolxemi Putzeys, 1875 Japan:Honshu(Tochigi:Chuzenji Lake nr.Nikko, "N.Nip."-LT); Sado Island
=decurtatus Lapouge, 1922 Chiuzenji nr.Nikko
=noesskei Emden, 1932 Sado Island


BACK



EUCARABUS Géhin, 1885; type species Carabus ulrichi Germar
species: 6



1.catenulatus Scopoli, 1763 Slovenia, Croatia, Bosnia, NE-Italy
ssp. catenulatus Scopoli, 1763 Slovenia("Kranj"); Croatia, Bosnia; NE-Italy:Trieste env., SE-Alps; Switzerland:Lugano env.
=catenatus Panzer, 1805 "Kranj"
=herbsti Dejean, 1826 "Mts.of Croatia"(alpine form)
=alternatus Haury, 1855
=subvirescens Motschulsky, 1865 "Idria"
~atropurpureus Dalla Torre, 1877; un
~atrovirescens Dalla Torre, 1877; un
~purpurescens Dalla Torre, 1877; un
~virescens Dalla Torre, 1877; un
=korlevici Hoffmann, 1883 Croatia:"Lic plateau"
=albanensis Géhin, 1885
~subvirens Haury, 1885; iss of subvirescens Motschulsky
=troyeri Born, 1911 Croatia:Gospic
~aenescens Depoli, 1929; un
=hosti Depoli, 1929 "Fiume env."(Rijeka)
=rattii Mandl, 1966 Mt.Palanzone nr.Lago de Como(=ca.20km NE Como)

ssp. fontanellae Reitter, 1896 Croatia:Adriatic Sea coast & islands; "Zara"(=Zadar)
=dinaricus Apfelbeck, 1904 Dinara Mts.:Troglav, Kamesnica
=mosorensis Müller, 1930 "Mosor"(=nr.Split)



2.cumanus Fischer, 1823 C,W-Caucasus; S-Georgia; S-Eur.Russia:Stavropol; "Desertus Cumanus"
=sobrinus Ménétriés, 1832
=submicans Reitter, 1888 "Circassia"
=laetulus Reitter, 1896 "Circassia"
~lagodai Lutshnik, 1911; un Terek River



3.italicus Dejean, 1826 N,C-Italy, S-Switzerland

ssp. italicus Dejean, 1826 N-Italy(Piemonte), S-Switzerland
=halbherri Lapouge, 1898
=ronchettii Born, 1900 Lombardia:Val Sassina
~battonianus Mandl, 1966; un Liguria:Pra:Mt.Riondo

ssp. rostagnoi Luigioni, 1904 C-Italy(Latium:Maccarese, Cisterno)
~rostagnonis Seidlitz, 1904; iss of rostagnoi Luigioni



4.obsoletus Sturm, 1815, nec Olivier, 1795; nbm Carpathians

ssp. obsoletus Sturm, 1815, nec Olivier, 1795; nbm Carpathians:Mor., Slov., SE-Poland, N-Hungary, E-Ukr., N,C-Rom.; >"Österreich"(=?Moravia)
=sacheri Thomson, 1875 ?"Beskiden"
=aureocupreus Reitter, 1880 "Trencsen"(=CW-Slovakia:Trenèín)
=csikii Mallasz, 1901 "Kom.Bestercze-Naszod"(=NC-Romania:Bistrita-Nasaud)
=prunneri Mallasz, 1901 "Kom.Also-Fehér"(=Alba Iulia)
~mallaszi Lapouge, 1908; un "N-Hungary"(=currently NW-Romania)
~uhligi Lapouge, 1908; un
=uhligi Holdhaus, 1910 "Ceahlau, Caliman, Rareul, Nagy-Hagymas"(=Rarau, Haºmaºul Mare)
=dudichi Csiki, 1927; nom.pro sacheri Thomson ?"Beskiden"
~bicoloratus Fleischer, 1930; un
~mallaszianus Breuning, 1932; un; nom.pro uhligi Holdhaus
~tesari Niedl, 1957; un M-Ukraine:Mukachevo

ssp. carpathicus Palliardi, 1825 C,SW-Romania:Fagaraº Mt.R. to Banat; "Transsylvanian Alps"
=euchromus Palliardi, 1825 "Banat"
=bielzi Birthler, 1886 C-Romania:"Çibin"(Çindrel Mt.R.)
=deubelianus Fleischer, 1911 "Kapellenberg nr.Kronstadt"(=Braºov)

ssp. nagyagensis Seidlitz, 1888; nom.pro procerus Bielz W-Romania:Bihor Mts.; Cluj; Hunedoara & Alba:("Nagyag, >Zalatna"=Sacarimb, Zlatna)
~carpathicusnagyagensis Birthler, 1886; un "Nagyag, Zalatna"(=Sacarimb, Zlatna)
~procerus Bielz, 1887, nec Reitt.; nom.pro carpathicusnagyagensis Birth. "Nagyag, Zalatna"(=Sacarimb, Zlatna)
=fossulifer Fleischer, 1893 "Kom.Bihar:Hadymadfalva"
~cupreus Fleischer, 1930; un
~viridis Fleischer, 1930; un
=tippmanni Breuning, 1936 "Zalatna"(=Alba:Zlatna)
~paranagyagensis Lie, 1990; un Arad/Hunedoara:Savirsin, Pojoga



5.parreyssi Palliardi, 1825;Croatia, Bosnia; Montenegro
ssp. parreyssi Palliardi, 1825; ?iss of paraysi Palliardi Croatia, Bosnia; Montenegro:Volujak, Durmitor Massif
=gattereri Géhin, 1882 "Steiermark"(error)
=purpurifer Reitter, 1885
=dobiaschi Beuthin, 1894 "Bosnia"
=padewiethi Csiki, 1906 S-Velebit Mts.:Stirovaca
~ganglbaueri Apfelbeck, 1890, nec Reitter, 1888 S-Bosnia:?
=planinaecola Csiki, 1906; nom.pro ganglbaueri Apfelbeck S-Bosnia:?
=squamisculptus Reitter, 1908; nom.pro ganglbaueri Apfelbeck S-Bosnia:?

ssp. plassensis Born, 1907 Bosnia:Adriatic Sea coast(Plasa); E-Serbia:Frushka Gora
=strauchi Born, 1908 N-Bosnia:Bihac env.
=tippmannianus Breuning, 1943 E-Serbia:Frushka Gora



6.ulrichi Germar, 1824; C,E-Europe, Hungary, Ukraine, Romania, Bulgaria, N-Yugoslavia


ssp. ulrichi Germar, 1824; ?iss of ulrichii Germar CE-Germany, S-Poland, Bohemia to Slovakia, NE-Hung., Ukraine; N,C-Romania; "Schlesien"
~ulrichii Germar, 1824; ?cos of ulrichi Germar
=leuckarti Petri, 1885 "Siebenburgen:Schässburg"(=Sighiºoara)
=podolicus Semenov, 1898 "Vinnitza"(=W-Ukraine:Vinnitsa)
=comanensis Born, 1902 SE-Romania:Dobrogea lowland:Comana, Braila; NE-Bulgaria:Razgrad
=pavlitscheki Born, 1907 "Bukowina"
=tigancanus Born, 1908 SE-Romania:Dobrogea lowland:"Tiganca"
=intercessor Sokolar, 1912 NC-Hungary:Nógrád:Rétság
~kardaschi Fleischer, 1926; un "Vlarapass"(=Moravia:Vlárský prùsmyk)
=volhynicus Lutshnik, 1927 "Wolhynia"
=planitiae Csiki, 1929 E-Hungary(Alföld region:?Debrecen, ?Gyula), Romania
=donovalensis Smetana, 1951 C-Slovakia:Nízké Tatry Mt.R.:Donovaly

ssp. rhilensis Kraatz, 1876 Austria, Belg., Holl., Germany, W-Bohemia, Hungary, CW-Romania, N-Yug.; Bulgaria(Rila Mts.)
~cupreoaeneus Dalla Torre, 1877; un "Oberösterreich"
~picipes Dalla Torre, 1877; un
~viridiaeneus Dalla Torre, 1877; un
=viridulus Kraatz, 1878
~parvus Géhin, 1885; un "Steiermark"
~stussineri Géhin, 1885; un Slovenia:"Krain"
=jaroslawi Fleischer, 1898 "Kom.Bihar"
=werneri Reitter, 1899 Bosnia:Vradac
=papukensis Koca, 1900 Serbia:Slavonia:Papuk Mts.
=pernix Csiki, 1904
=slivensis Apfelbeck, 1904 CE-Bulgaria:Sliven
=sokolari Born, 1904 S-Austria:Leitha Mts.
=baranyensis Sokolar, 1908 SC-Hungary:"Kom.Baranya"
~germanicus Sokolar, 1908, nec Semenov, 1898 "Germany, Nieder- & Oberösterreich"
=alamannicus Csiki, 1927; nom.pro germanicus Sokolar "Germany, Nieder- & Oberösterreich"
~stegmanni Mandl, 1969; un Niederbayern:Straubing
~serbomacedonicus Kenyery, 1970; un S-Serbia:?
~kucajensis Paviæeviæ & Toševski, 1988; un NE-Serbia:Kuèaj Mts.:Babina Glava:above Resavska peèina Cave
=nastasi Paviæeviæ, 1995 Serbia:?

ssp. fastuosus Palliardi, 1825 SW-Romania:Banat(Mehadia)
~cupreonitens Kraatz, 1878, nec Chevrolat, 1861
~parallelus Kraatz, 1878, nec Faldermann, 1835 "Banat:Nemet Bogsan"(=Bocsa nr.Reºita)
~robustus Kraatz, 1878, nec Deyrolle, 1869 "Banat"
~superbus Kraatz, 1878, nec Schaufuss, 1872 "Banat:Moldowa"(=Moldova Noua)
~viridilimbatus Kraatz, 1878, nec Motschulsky, 1859 "Banat:Nemet Bogsan"(=Bocsa nr.Reºita)
=glaucus Haury, 1878 Banat:Bazias
~cyaneus Bodemeyer, 1881, nec Fabricius, 1776 Banat:Reºita
~cuprinus Lapouge, 1902; un Banat:Bazias
~fabrizioi Battoni & Breuning, 1970; un E-Serbia:Negotin
~transdiernae Paviæeviæ & Toševski, 1988; un NE-Serbia:Tekija(=Dunav right bank opposite to Orºova)
~alionidiernae Lie, 1989; ?nud Banat:Alion nr.Orºova
~superboides Lie, 1989; ?nud Banat:"sanatoriul Marilla de linga Oravita"(=Oravita env.)
~alioni Lie, 1990; un; ?nom.pro alionidiernae Lie Banat:Alion nr.Orºova
~ciutagrosi Lie, 1990; un Banat:dealul Ciuta nr.Groºi(=ca.15km NE Faget)
~feregarii Lie, 1990; un Banat:Feregari/Domogled valley nr.Baile Herculane
~subfastuosus Lie, 1990; un Banat: betw.Cavaran/Tincova(=ca.30km SE Lugoj)
~subsuperbus Lie, 1990; un; ?nom.pro superboides Lie Banat:"sanatoriul Marilla de linga Oravita"(=Oravita env.)
~topeniae Savulescu, 1992; un Banat:Topenia valley(=NE of Baile Herculane)
=subfastuosus Lie, 1996 Banat: betw.Cavaran/Tincova(=ca.30km SE Lugoj)
~subsuperbussuperboides Lie, 1996; un Banat:"sanatoriul Marilla de linga Oravita"(=Oravita env.)
~umbrosus Lie, 1996; un Banat:Poiana Rusca Mts.:Nadrag & Borlova env.

ssp. arrogans Schaum, 1858 NE-Serbia:Majdanpek, Donji Milanovac; "Serbia"
~ceremosnjensis Paviæeviæ & Toševski, 1988; un NE-Serbia:Homolje Mts.:Ceremošnja
~gornjakensis Paviæeviæ & Toševski, 1988; un NE-Serbia:Šetonje:Gornjak monastery env.
~pseudoarrogans Paviæeviæ & Toševski, 1988; un NE-Serbia:Donji Milanovac


BACK



MORPHOCARABUS Géhin, 1885; type species Carabus monilis Fabricius
species: 20



1.aeruginosus Fischer, 1822 E-Eur.Russia, Siberia, Altays, Sayans

ssp. aeruginosus Fischer, 1822 E-Eur.Russia, Siberia, Altays("Ridders"=Leninogorsk; ?by error), Sayans
=aereus Dejean, 1826
=gmelini Fischer, 1827
=incertus Motschulsky, 1846 "Sib.or."(LT)
=subcostatus Motschulsky, 1850 "Sib.or."
~capucinus Géhin, 1885; un
=cereus Lapouge, 1912 "Transbaykalia"

sp. hermani Mannerheim, 1827 Urals(Zlatoust), W-Siberia

ssp. salechardensis Obydov, 1999 NW-Siberia(Salekhard env.) & NE-Eur.Russia; ?NC-Siberia

ssp. eschscholtzi Mannerheim, 1827 Altays, Sayans, N-Mongolia; "Sibiria"

ssp. tulatensis O.Berlov & Shilenkov, 1996 Altays:Tigiretsky Mt.R.(Tulata env.)

ssp. zyrjanovskianus Shilenkov & O.Berlov, 1996 W-Altays:Bukhtarma River basin(Zyryanovsk)

ssp. verzhutzkii O.Berlov & Shilenkov, 1996 NW-Altays:Anuysky & Seminsky Mt.R.:Ch’ornyy Anuy env., Shebalino env.

ssp. kolyvanus Shilenkov & O.Berlov, 1996 Russia:NW-Altays:Kolyvansky Mt.R.:Kolyvan env.:Kamenka village



2.chaudoiri Gebler, 1847 C, NE-Siberia, ?N-Mongolia

ssp. chaudoiri Gebler, 1847 Mts.nr.Kuznetsk(=Kuznetsky Alatau) & Kan River(=E of Krasnoyarsk); "Altay"(LT)

ssp. mongolorum Csiki, 1927; nom.pro neglectus Lapouge, 1913 N-Mongolia; "Tunkun, Sajan"(=Siberia/Mongolia:Tunkunskye Goltsy Mts.; LT)
~neglectus Lapouge, 1913, nec Kraatz, 1887 "Tunkun, Sajan"(=Siberia/Mongolia:Tunkunskye Goltsy Mts.; LT)

ssp. czadanicus Obydov, 1997 W-Tuva:Chadan(error; correctly:C-Tuva:Khadyn ca.50km NW Kyzyl)

ssp. marusiki Obydov, 1997 SE-Tuva:Sangilen Mts.:Kargy River valley:50:31N/97:01E

ssp. zaikai Obydov, 1999 C-Tuva:Akademika Obrucheva Mt.R.:Sainak Pass, 52:00N/95:34E

ssp. beladici Obydov, 2001 Tuva:East Tannu-Ola Mt.R.:Chongyz Tayga, 1500m, Kara-Kohl Lake

ssp. juliae Obydov, 2002 Tuva:Mungun-Tayga:Khindiktig-Khol Lake, 2400m

ssp. shilenkovi O.Berlov, 1989Tuva to NE-Siberia(Chukot Peninsula:Bilibinsk:Omolon vill.); Yakutsk:Ust’-Yansk



3.excellens Fabricius, 1798 NE-Romania, Moldavia, Ukraine; W-Russia: Kursk env.; Byelorussia
=goldeggi Duftschmid, 1812 "Podolia"
=erythromerus Dejean, 1826 Moldavia:Bendery
=erythrocnemus Fischer, 1827
=erythrodes Fischer, 1827
~chrysitis Letzner, 1850; un
~polonicus Lomnicki, 1886, nec Motschulsky, 1850 "Galicia:Lubaczow"(=SE-Poland:Lubaczow E of Rzeszów)
=frivaldskyi Kraatz, 1887 "Galicia"
=lomnitzkii Reitter, 1896; nom.pro polonicus Lomnicki "Galicia:Lubaczow"(=SE-Poland:Lubaczow E of Rzeszów)
=sturmi Schilsky, 1899
=moldaviensis Born, 1903 S-Moldavia
=jasilkowskii Born, 1905 "Moldavia:Bojan"
~pulchellus Csiki, 1906, nec Brüggemann, 1873 "Tavarnók"(=SW-Slovakia:Tovarníky; error)
~indiconotus Lapouge, 1906; un
~paradoxus Lapouge, 1906; un
~rubicundus Lapouge, 1906; un
~pseudomoldaviensis Hormuzaki, 1907; un NE-Romania:"Jassy"(=Iaºi)
=pyretanus Hormuzaki, 1907 N-Moldavia
=rareulensis Born, 1907 NE-Romania:Rarau Mts.(alpine form)
~paradoxalis Lapouge, 1909; un
=feketei Csiki, 1926; nom.pro pulchellus Csiki "Tavarnók"(=SW-Slovakia:Tovarníky; error)
~botezati Marcu, 1934; un
~striatulus Marcu, 1934; un



4.gebleri Fischer, 1817 Altays Mts.:Zmeinogorsk, Zyryanovsk env.

ssp. gebleri Fischer, 1817 Altays Mts.:Zmeinogorsk env.

ssp. ultimus Obydov, 1999 Altays Mts.:E-Kazakhstan:Bukhtarma River valley(Zyryanovsk env.)


5.henningi Fischer, 1817E-Eur.Russia, Urals; C,W-Siberia, Altays, Sayans; ?N-Mongolia

ssp. henningi Fischer, 1817 E-Eur.Russia, Urals, W-Siberia(Barnaul), Altays, Sayans
=sahlbergi Dejean, 1826 Altays, Sayans, N-Mongolia, Baykal Lake env.; "Siberia"
~ruficornis Kraatz, 1882, nec DeGeer, 1774 Altays
=spurius Kraatz, 1882
~unicolor Kraatz, 1882, nec Fabricius, 1787
=continuus Lapouge, 1906"Jelan"(=Baykal Lake env.)
=roeschkei Csiki, 1901 Yenisey River
=sajanicus Csiki, 1901 Sayans
=barnaulanus Born, 1922 Barnaul
=uralicus Born, 1922 Urals

ssp. oviformis Beheim & Breuning, 1943 arctic W-Siberia("Kantaika"=Khantayka SW of Noril’sk)

ssp. hamarensis Shilenkov, 1996 Russia:Hamar-Daban Mt.R.:Snezhnaya River valley

ssp. melyachi Shilenkov & Menstshikov, 1996 Russia:W-Altays:(Kolyvansky Mt.R.:Kolyvan env.:Kamenka village)

ssp. saldaitisi Obydov, 1997 SE-Tuva:Sangilen Mts.:Balyktyk-Khem River source:50:18N/96:24E, 2400m

ssp. peczoricus Obydov, 2000 NW-Siberia:Komi:Usa:Pechora basin:Kosiyu River mouth, Lek-Rogovaia River mouth


6.hummeli Fischer, 1823Siberia; Altays, Sayans, Mongolia, Far East, Korea, NE-China

ssp. hummeli Fischer, 1823 C,W,E-Siberia(Nerchinsk); Altays, Sayans, Mongolia
=burnaschewi Dejean, 1826 "Siberia"
=cyaneoviolaceus Motschulsky, 1844 "Dahuria, Irkutsk"
=obversus Motschulsky, 1846 Nerchinsk(LT)
=ochoticus Mannerheim, 1852 Far East:"Port Ajan"
~multistriatus Lapouge, 1924, nec Motschulsky, 1865
=densestriatus Csiki, 1927; nom.pro multistriatus Lapouge

ssp. middendorfi Ménétriés, 1851Russia:Far East:Bolshoy Shantar Island

ssp. stolidus Lapouge, 1924?SE-Siberia:Hamar-Daban Mt.R.; ?Sayans

ssp. jurgae Obydov, 1995 Yakutsk" Suntar-Khayata Mt.R.: Khandyga R.(error; in fact: tundra 300km N Khandyga R.)

ssp. nevelskii Shilenkov, 1996 S-Sakhalin(Kril’on Peninsula:Kirilovo); C-Sakhalin

ssp. smaragdulus Kraatz, 1878, nec Fabricius; 1798; nbm Russia:Far East(middle Amur River), N-Korea, NE-China:Jilin
=chrysothorax Kraatz, 1886 Suyfun River(=W of Vladivostok)
~decoloratus Kraatz, 1886, nec Reitter, 1884 Suyfun River(=W of Vladivostok)
=purpuricollis Kraatz, 1886 Suyfun River(=W of Vladivostok)
=purpuripennis Kraatz, 1886 "middle Amur River"
=subaeneus Kraatz, 1886 Suyfun River(=W of Vladivostok)
=subauratus Kraatz, 1886 Suyfun River(=W of Vladivostok)
=viridimarginatus Kraatz, 1886 Suyfun River(=W of Vladivostok)
~viridipennis Kraatz, 1886, nec Lewis, 1880 "Amur"
=buriatus Lapouge, 1924 "middle Amur River"
=amurlandicus Shilenkov, 1994; nom.pro viridimarginatus Kraatz Suyfun River(=W of Vladivostok)

ssp. tristiculus Kraatz, 1879 Far East(Suyfun River mouth W of Vladivostok)
=gracilentus Kraatz, 1881 "Innshan Mts.W of Peking"

ssp. pusongensis Imura, 1993 NC-Korea:Nangnim Mts.:Pusong, Paektu-san Mass.; NE-China:Jilin; Far East:Vladivostok env.
=gustavi Shilenkov, 1996 Far East:Vladivostok env.:Chornye Gory Mts.:Kedrovaya Pad’
=wangxingensis Deuve & Li, 1998 E-Jilin:"Wangxing Xian:Daxinggou"(correctly:Wangqing Xian)

ssp. suensoni Mandl, 1979 NE-Shanxi:NE of Xin Xian:Wutai Shan Mts.; Fang Shan W of Taiyuan
~fuscipennis Mandl, 1979; un NE-Shanxi:NE of Xin Xian:Wutai Shan Mts.



7.karpinskii Kryzhanovskij & Matveev, 1993 SW-Siberia:South Urals:Mt.Iremel, Yamantau, Zigalga



8.kozhantschikovi Lutshnik, 1924 West Sayans:Kazyr Suk River
=martjanovi Lutshnik, 1924 West Sayans:Kazyr Suk River



9.mestscherjakovi Lutshnik, 1924 West Sayans, East Sayans, E-Altays

ssp. mestscherjakovi Lutshnik, 1924 West Sayans(Buyba Lake), East Sayans

ssp. pseudoodoratus Brinev, 2002 Russia:Krasnoyarsk:Us River:7km SW Aradan vill.

ssp. zherichini Shilenkov, 1990 N-Siberia:Polar Urals to Kolyma River(Gydan Peninsula:Nosok village)
=kuzminae Shilenkov, 1990 NE-Siberia:Kolyma basin:Kolyma Lowland:middle Alazeya River
=pjasinensis Gottwald, 1993 NC-Siberia:Taymyr Peninsula:Pyasina River
~taimyrensis Shilenkov, in litt.

ssp. shaman Shilenkov, 1996 Russia:West Sayans:Mt.Shaman
=sambylensis Shilenkov, 1996 West Sayans:Dzhembashsky Mt.R.:Abakan River basin:Sambyl Pass

ssp. telezkianus Shilenkov, 1996 Russia:E-Altays:Teletskoe Lake env.:(SW-bank:Mt.Altyntu); Chulyshman Mts.
=altyntuanus Shilenkov, 1996 Russia:E-Altays:Chulyshman River:Mt.Altyntu

ssp. tuvinicus Obydov, 2002 Russia:Tuva:Uyuk River valley:Taskyl Mt.R.; 1f



10.michailovi Kabak, 1992 NE-Kazakhstan:Sarym-Sakty Mt.R.:45km WSW Katon-Karagay:E of Ornek



11.monilis Fabricius, 1792France, Belgium, Holland, Switzerland, Germany(Halle), Brit.Isles, Ireland
~affinis Panzer, 1798, nec Schrank, 1781 Germany:Würzburg
=consitus Panzer, 1809 Bern
=kronii Hoppe & Hornschuch, 1825 Salzburg(error) & Mühlhausen
=amoenus Baudet-Lafarge, 1836 Puy-de-Dome:Mt.Doré
=?helveticus Heer, 1837; ?hyb Switzerland:?"Guttannen, Herzogenbuchsee"; ?hyb monilis/cancellatus
=schartowi Heer, 1838 "Jura"
=gracilis Küster, 1846 "Bayern"
~regularis Wissmann, 1848, nec Fischer, 1823 "Thüringen"
=scolecaster Gistl, 1857 "Helvetia"
=veris Gistl, 1857 "Germany"
=rufofemoratus Motschulsky, 1865 "Helvetia"
=taunicus Heyden, 1871 W-Germany:"Mt.Taunus & Vogelsberg"(alpine form)
~femoralis Géhin, 1876, nec Motschulsky, 1865 ~azurescens Dalla Torre, 1877; un
~viridescens Dalla Torre, 1877; un
=alticola Bellier, 1880 Alpes-Hte-Provence:Digne env.
=anomalus Géhin, 1880 Switzerland:Mt.Pilatus
=femoratus Géhin, 1880
=interpositus Géhin, 1880 "Parigi"
=rugatinus Géhin, 1880
=sabaudus Géhin, 1880 "Jura"
~nigritulus Kraatz, 1881, nec Kraatz, 1876 "Grande Chartreuse"
~rubricrus Géhin, 1885; un; nom.pro rufofemoratus Motschulsky "Helvetia"
~trilineatus Haller, 1885, nec Fabricius, 1792 "Schweizer Jura"
~varicolor Joerin-Gerber, 1888; un "Jura"
=bavaricus Beuthin, 1896 Bayern:Lechfeld
~interruptus Beuthin, 1896, nec Fabricius, 1775
=moestulus Beuthin, 1896
=sengstacki Beuthin, 1896
=?subschartowi Lapouge, 1898; ?hyb Haute-Garonne:Luchon
=meridionalis Born, 1905 S-France
~anglicus Lapouge, 1906; un England:?
~externus Lapouge, 1906; un
~gaultieri Lapouge, 1906; un Loire-Atlantique:Nantes
=tugenus Lapouge, 1906 "Kanton Appenzell"
=perreti Born, 1907 Isere:Notre-Dame-la-Salette(alpine form)
~arvernus Barthe, 1908; un Puy-de-Dome:Mt.Doré
~insularis Born, 1908, nec Hope, 1837 England:?
~bavariensis Lapouge, 1909; iss of bavaricus Beuthin
=pustulatus Lapouge, 1909 Puy-de-Dome
=tigurinus Lapouge, 1909
~kellneri Hubenthal, 1910; un "Thüringen"
=nicodii Pic, 1910
=rhodanicus Lapouge, 1916 S-France:?
=sequanus Lapouge, 1916 Cote-d’Or:Brazay en Morvan
=succisus Lapouge, 1916 France:St.Gaudens
=subpyrenaeus Barthe, 1924 Haute-Garonne:St.Gaudens, Col d’Aspin
~charbonieri Sirguey, 1931; un Puy-de-Dome:Mt.Doré
~cylindricus Sirguey, 1931, nec Herbst, 1784 Puy-de-Dome
~cylindricusniger Sirguey, 1931; un Puy-de-Dome
~cylindricusviolaceus Sirguey, 1931; un Puy-de-Dome
~femoratusviolaceus Sirguey, 1931; un
=incostatus Sirguey, 1931 Puy-de-Dome
~incostatusfemoralis Sirguey, 1931; un Puy-de-Dome
~incostatusniger Sirguey, 1931; un Puy-de-Dome
~incostatusviolaceus Sirguey, 1931; un Puy-de-Dome
~interposituscyaneus Sirguey, 1931; un
~interpositusniger Sirguey, 1931; un
~interpositusviolaceus Sirguey, 1931; un
=lavagnei Sirguey, 1931 Puy-de-Dome:Mt.Doré
~marcoti Sirguey, 1931; un
~pustulatusniger Sirguey, 1931; un ?Puy-de-Dome
~pustulatusviolaceus Sirguey, 1931; un ?Puy-de-Dome
~rhodanicusviolaceus Sirguey, 1931; un
~sequanusniger Sirguey, 1931; un S-France:?
~sequanusviolaceus Sirguey, 1931; un S-France:?
~theroudi Sirguey, 1931; un
=theryi Sirguey, 1931
~trapeti Sirguey, 1931, nec Bleuse, 1885 Puy-de-Dome:Mt.Doré
~trilineatusniger Sirguey, 1931; un ?"Schweizer Jura"
~trilineatusviolaceus Sirguey, 1931; un ?"Schweizer Jura"
~saouensis Mandl, 1967; un Drôme:Ft.de Saou
=daniellae (Bellon & Tarrier, 1972) ?Alpes-Maritimes:Col de Champs
=ventusicus (Bellon & Tarrier, 1972)
=montichares (Hansen & Gaskowiak, 1973) Ariége:Pailhéres
=dupeuxi Deuve, 1974 Charente:Foret de Braconne
=benoiti (Devecis, 1975) Corréze:Albussac
~meneensis Grotz & Grotz, 1975; un Drôme:Col de Menée
=norensis Pham, 1977 Aude:Pic de Nore
=rosayanus Dufour, 1978 Cote-d’Or:Rosay
=fagnensis Grotz, 1978
~aoustensis Machard, 1979; un Drome:Aouste
~brunetensis Machard, 1979; un Plomb du Cantal:Prat de Bouc
~marsannensis Machard, 1979; un Drome:Ft. de Marsanne
~ourthizetensis Machard, 1979; un E-Pyrenees:Pic d’Ourthizet
~pailherensis Machard, 1979; un Ariége:Port de Pailhéres
~proncelensis Machard, 1979; un Vercors Mts.:Col de Proncel
~nepos Ivanovs, 1981; un N-Germany:Niedersachsen:Verden an der Aller(=SE of Bremen);
=?excellens Fabricius
=alpigradus Sermet, 1985 S-Switzerland:Valais:Saas valley
~vincentianus Devecis, 1991; un Cantal:Col d’Aulac
~barilloti Devecis, 2000; un Aveyron:NE-part:Plateau d’Aubrac(HT not designated)



12.odoratus Motschulsky, 1844; ?non priority Siberia, Altays, Sayans; ?N-Mongolia

ssp. odoratus Motschulsky, 1844 Russia:Baykal Lake:SE-coast; Hamar-Daban Mts.(LT)
~vulneratus Lapouge, 1915; un =batesianus Deuve, 1995 "Baikal"; 1m

ssp. viridilimbatus Motschulsky, 1845 SE-Siberia & Far East:Barguzin Mt.R., Amur; CE-Siberia:S of Yakutsk:Amga; "Amur bor."(LT)

ssp. irkoutskensis Lapouge, 1915 Irkutsk env.

ssp. dohrni Gebler, 1847; ?non priority C-Altays; "Siberia, Altay"(LT)
=?microchondrus Fischer, 1842; inc; ?priority "Songoria:Kreis Ajagus"(=?Ayaguz 300 km S Semipalatinsk)
~interstitialis Motschulsky, 1845; nud; nom.pro alysidotus: Fischer (Siberia:Irtysh River; invalid LT)
~irkoutskensis Géhin, 1885; nud; nom.pro interstitialis Motschulsky

ssp. baeri Ménétriés, 1851 NC,NE-Siberia:Yenisey to Kolyma River; "Boganida"(=S-part of Taymyr Peninsula)
=tolli Poppius, 1910 Chara-Ullah Mts.betw.Lena/Jana River(=Verkhoyansky Mt.R.)

ssp. chaffanjoni Lesne, 1897 arctic E-Siberia:Bulun nr.Lena River mouth

ssp. melleus Lapouge, 1909; ?un East Sayans:Mondy; ?NC-Mongolia

ssp. septentrionalis Breuning, 1932 NW,NC-Siberia:arctic Urals("Obdorsk"=Salekhard) to Yenisey River

ssp. kamtschatensis Breuning, 1942 NE-Siberia:Kamchatka:Penzhinskaya Guba:Shestakovo

ssp. antropovi Shilenkov, 1996 Russia:East & West Tannu-Ola Mt.R.(East Tannu-Ola:Shuurmak env.)

ssp. bargusinicus Shilenkov, 1996 Russia:Barguzinsky Mt.R.(Davshe River valley)

ssp. dabanensis Shilenkov, 1996 Russia:Irkutsk:Hamar-Daban Mt.R.:S-slope(Snezhnaya River valley)

ssp. krugeri Obydov, 1999 C-Tuva:Akad.Obrucheva Mt.R.:Koptu Pass, 51:56N/95:32E; Sainak P., 52:00N/95:34E

ssp. magadanicus Obydov, 1999 E-Siberia:Magadan env.(Magadan)

ssp. martjanovianus Obydov, 1999 Russia:West Sayans:Sayanogorsk

ssp. taskylensis Obydov, 2001 Russia:Tuva:Taskyl Mt.R.:Uyuk River, 60km NE Tuzan

ssp. jacuticus Obydov, 2002 Russia:Yakutia:Vostochaya Khandyga River:Teplyi Klyuch

ssp. putoranicus Obydov, 2002 Russia:NE-Siberia:Putorana plateau:Ajan Lake env.

ssp. chlebnikovae Obydov 2005; nom. pro marinae Obydov Russia:Severo-Baykalsk env.(80km S Severo-Baykalsk:5km S Kurkula R. mouth)
~marinae Obydov, 2003, nec Maux, 1995 Russia:Severo-Baykalsk env.(80km S Severo-Baykalsk:5km S Kurkula R. mouth)

ssp. czernyscheviellus Obydov, 2002 Russia:Chita:Nerchinsko-Kuengsky Mt.R.:Vershino-Darasunsky vill.env.:Mt.Bogatui



13.regalis Fischer, 1822 E-Eur.Russia; C,W-Siberia, Altays, Sayans; N-Mongolia

ssp. regalis Fischer, 1822 E-Eur.Russia; W-Siberia(Barnaul)
=cuprinus Fischer, 1822
=cyanicollis Dejean, 1826
=pasianax Fischer, 1827
=interstinctus Motschulsky, 1865 Irtysh River
~viridicinctus Kraatz, 1882, nec Schaufuss, 1872
=viridicollis Kraatz, 1882
=jureceki Born, 1922 "Mts.nr.Simbirsk"(=Ulyanovsk)
=volgensis Csiki, 1927; nom.pro jureceki Born "Mts.nr.Simbirsk"(=Ulyanovsk)

ssp. jenissoni Dejean, 1831 Altays, Sayans, C-Siberia, N-Mongolia
~dubius Motschulsky, 1846, nec Herbst, 1779 Irkutsk(LT)
=jacutus Mäklin, 1881 "Potapowski"(=Potapovo SE of Noril’sk)
~cyanescens Kraatz, 1882, nec Sturm, 1815 ?"Altay"
~nigritulus Kraatz, 1882, nec Kraatz, 1876 ?"Altay"
=fraudator Lapouge, 1906 Irkutsk
=hunnorum Lapouge, 1909 "Dzhungaria"

ssp. tschusovae Obydov, 2004 E-Kazakhstan:Ivanovsky Mt.R.(upper level of "Prokhodnoy Belok")



14.rothi Dejean, 1829 Romania, NE-Hungaria, W-Ukraine

ssp. rothi Dejean, 1829 C-Romania:Sibiu env.; "Transsylvania"
=aequistriatus Kraatz, 1878
=latestriatus Kraatz, 1878
=quadricatenatus Kraatz, 1878; nec ?
=varistriatus Kraatz, 1878
=birthleri Reitter, 1885
~rugulosus Birthler, 1885; un
~ignoscus Csiki, 1906; un; nom.pro quadricatenatus Kraatz

ssp. comptus Dejean, 1831 CW-Romania:Retezat Mts.to Bihor Mts.; "Banat"
~hopffgarteni Kraatz, 1880, nec Kraatz, 1878 "S-Hungary"(inexact; ?Godeanu & Tarcu Mts.)
~merkli Kraatz, 1880, nec merklii Hopffgarten, 1878 ~merklianus Géhin, 1885; un; nom.pro merkli Kraatz
=dieneri Merkl, 1900 "Szöreny Mts."(=Godeanu/Tarcu Mts.)
=diffinis Csiki, 1906 "Komitat Kolosz"(=Cluj)
=szoerenyensis Csiki, 1908; nom.pro hopffgarteni Kraatz "S-Hungary"(inexact; ?Godeanu/Tarcu Mts.)
~ulrichhofmanni Kleinfeld & Korell, 1986; un SW-Romania:30km SE Lugoj:Cavaran
~cavarani Lie, 1989; un SW-Romania:30km SE Lugoj:betw.Cavaran/Tincova
=ulrichhoffmanni Lie, 1989 SW-Romania:ca.20km E Lugoj:Nadrag env.:Gosta Hut nr.Nadrag
~borlovensis Lie, 1994; un SW-Romania:ca.50km SE Lugoj:Mt.Muntele Mic near Borlova
~micotensis Lie, 1999; un SW-Romania:ca.40km SE Lugoj:Poiana Rusca massif:Micota valley nr.Boutari

ssp. hampei Küster, 1846 NW,C,?SW-Romania; NE-Hung.:Szabolcz-Szatmár; W-Ukraine:Mukachevo; "Siebenburgen"
~schaumi Birthler, 1886, nec Gaubil, 1849 "Serbia"(?inexact)
=aurosericeus Kraatz, 1880 "Maros-Vasarhely"(=Tirgu Mures)
=marusii Ormay, 1890 "Gyula-Fehervar & Nagy Enyed"(=Alba Iulia & Aiud)
=ormayi Reitter, 1896 "Beregszasz"(=Beregovo)
=liebli Dietl, 1897 "Komitat Kis-Küküllö"(=Tirnava Mic)
~marginatus Kraatz, 1900; un ?"Maros-Vasarhely"(=Tirgu Mures)
=zoppai Kraatz, 1900 ?"Maros-Vasarhely"(=Tirgu Mures)
=dacicus Csiki, 1906 "Szamosujvar & Naszod"(=Gherla & Bistrita Nasaud)
=eximius Csiki, 1906 "Nagyvarad"(=Oradea)
=fraternus Csiki, 1906 "Maros-Vasarhely"(=Tirgu Mures)
~mendax Csiki, 1906, nec Rossi, 1790 "Komitat Marmaros"(=Maramureº)
~validus Csiki, 1906, nec Kraatz, 1884 "Komitat Hunyad:Nagayg"(=Hunedoara:ca.40km NE Deva:Sacarimb)
~vitiosus Csiki, 1906; un
~zilahiensis Csiki, 1906; un "Komitat Szilagy:Zilah"(=Salaj:Zalau)
~concinnus Lapouge, 1916; un; nom.pro schaumi Birthler "Serbia"(?inexact)
=hunyadensis Csiki, 1926; nom.pro validus Csiki "Komitat Hunyad:Nagayg"(=Hunedoara:ca.40km NE Deva:Sacarimb)
=bokori Csiki, 1927; nom.pro concinnus Lapouge "Serbia"(?inexact)
=telekii Csiki, 1937 ?"Banat"(?inexact)
=gutiiensis Takács & Lie, 1992 NW-Romania:NE of Baia Mare:Gutii Mts.
=praedacicus Takács & Lie, 1992 C-Romania:WNW of Alba Iulia:Zlatna

ssp. incompsus Kraatz, 1880 NC,C-Romania:Rodna Mts.to Harghita Mts.; "S-Hungary:Brasso env."(=Braºov)
~elegantissimus Csiki, 1906; un =mehelyanus Csiki, 1906 "Komitat Csik:Vigyazok-Tarko"(=ca.Harghita Mts.)
=spectabilis Csiki, 1906 Rodna Mts.

ssp. alutensis Savulescu, 1972 C-Romania:Olt River:Rimnicu Vilcea env.(Calimaneºti)



15.scheidleri Panzer, 1799 CE-Europe, W-Carpathians, Balcans

ssp. scheidleri Panzer, 1799 N,E-Austria(Wien); SE-Germany:Passau, Regensburg; S-Bohemia; S-Moravia; C,W-Hungary

~caeruleus Panzer, 1812, nec coeruleus Steven, 1808 Wien
=preyssleri Duftschmid, 1812 Niederösterrieich; Puchberg am Schneeberg(LT)
=aeneipennis Sturm, 1815
=purpuratus Sturm, 1815
~virens Sturm, 1815, nec Müller, 1776
~atrocoeruleus Letzner, 1850; un
~aureus Letzner, 1850; un
~cyaneus Letzner, 1850; un
~niger Letzner, 1850; un
~purpureoaureus Letzner, 1850; un
~purpureomarginatus Letzner, 1850; un
~purpureus Letzner, 1850; un
~viridiaeneus Letzner, 1850; un
~viridimarginatus Letzner, 1850; un
~viridis Letzner, 1850; un
=gibbicollis Motschulsky, 1865
=ambitiosus Schaufuss, 1872
=improbus Schaufuss, 1872
=principatus Schaufuss, 1872
=superbus Schaufuss, 1872
=viridicinctus Schaufuss, 1872
~aerescens Dalla Torre, 1877; un
~aeruginescens Dalla Torre, 1877; un
~aurescens Dalla Torre, 1877; un
~coerulescens Dalla Torre, 1877; un
~cuprescens Dalla Torre, 1877; un
~elegans Dalla Torre, 1877; un
~fuscescens Dalla Torre, 1877; un
~metallescens Dalla Torre, 1877; un
~nigrescens Dalla Torre, 1877; un
~violescens Dalla Torre, 1877; un
~virescens Dalla Torre, 1877; un
=dominus Reitter, 1885
~graciosus Géhin, 1885; un
=styriacus Kraatz, 1887 "Styria"(=Steiermark)
=limbifer Reitter, 1889
=burghauseri Fleischer, 1899
=floriani Penecke, 1905 Steiermark:Gratwein
~distinguendus Csiki, 1906, nec Duftschmid, 1812 E of Budapest:Rakosfeld
=jucundus Csiki, 1906 W of Budapest:Pilis Mts.
=pannonicus Csiki, 1906 C-Hungary:Fejér:Székesfehérvár
~duchoni Lapouge, 1906; un "Krain"(error)
~prolifer Lapouge, 1906; un "Krain"(?error)
=kollarisans Sokolar, 1907 "Moravia"
~coeruleocinctus Fleischer, 1925; un
~elegans Fleischer, 1925; un
=gebhardti Bodemeyer, 1927 SW-Hungary:Tolna:Dombovar
=maderianus Mandl, 1957 Niederösterrieich:Aspang
~pseudoscheidleri Mandl, 1964; un NE-Austria:Leitha Mts.:Winden env.
~punctatocostatus Mandl, 1964; un Leitha Mts.:Winden env.
~reductecostatus Mandl, 1964; un Leitha Mts.:Winden env.
~baderlei Mandl, 1965; un NE-Austria:Burgenland:Parndorfer Platte(SE of Wien)
~praescheidleri Mandl, 1965; un SW-Hungary:Baranya:Mescek Mts.
~pseudojucundus Retezár, 1974; un NW-Hungary:Komárom:Gerecse Mts.
~vertesensis Retezár, 1974; un NW-Hungary:Fejér:Vértes Mts.
=csikiellus Deuve, 2003; nom. pro distinguendus Csiki E of Budapest:Rakosfeld

ssp. helleri Ganglbauer, 1893; ?non priority Slovakia("Kom.Neutra"=Nitra); N,C,E-Moravia; N-Bohemia; ?W-Ukraine
=?repercussus Drapiez, 1819; ?priority "Karpathen"
~morawitzi Kraatz, 1887, nec Ganglbauer, 1887 ?"Bohemia"
~cupripennis Fleischer, 1925; un
~pseudoambitiosus Fleischer, 1925; un
~pseudoexcellens Fleischer, 1925; un
~pseudosuperbus Fleischer, 1925; un
~viridipennis Fleischer, 1925; un
~pseudopreyssleri Breuning, 1932; un; nom.pro morawitzi Kraatz ?"Bohemia"
=giuseppemuelleri Hildebrand, 1955 S-Moravia:"Brünn"(=Brno)
=beheimi Niedl, 1957 W-Ukraine:Mukachevo
~bieneri Mandl, 1965; un C-Slovakia:Kralovany
~marmaroschensis Mandl, 1965; un "Marmaros"
~subparvulus Mandl, 1965; un NC-Hungary:Börzsöny Mts.:Dios Jenö
~subtilepunctatus Mandl, 1965; un

ssp. kollari Palliardi, 1825 SW-Romania:Banat Mts.(Mehadia), Vilcan Mts.
=magnificus Kraatz, 1878
=semetricus Kraatz, 1878 "Banat"
=negotinensis Reitter, 1896 Serbia:Negotin
~paroecus Csiki, 1906; un
~sejunctus Csiki, 1906; un
=joergeri Born, 1927 "Walachei"

ssp. praecellens Palliardi, 1825 Serbia:Novi Sad, Fruska Gora Mts.("Futtak"=Futog)
~kollarioides Mandl, 1966; un Serbia:Fruska Gora Mts.

ssp. illigeri Dejean, 1826, nec Panzer, 1806; nbm Bosnia; W-Croatia:Dinara planina
=laticollis Sturm, 1826
=stentzi A.& J.B.Villa, 1833 "Hungary"(?error)
~parallelus Kraatz, 1878, nec Faldermann, 1835 Pljesevica Mts.
=curtulus Ganglbauer, 1892 Mts.nr.Sarajevo
=bjelasnicensis Apfelbeck, 1902 Bosnia:Bjelasnica Mts.
=koschanini Csiki, 1904 Serbia:Kopaonik Mts.
~transfuga Apfelbeck, 1907, nec Semenov, 1898 Sarajevo env.
=unanensis Born, 1909 Bosnia:Bihac
~cerensis (Toševski, 1990); un NW-Serbia:Cer Planina(=SW of Sabac), Korenita nr.Loznica
~tarensis (Toševski, 1990); un W-Serbia:Tara Mts.:Mitrovac

ssp. versicolor Frivaldsky, 1835 Bulgaria:Stara planina Mts.(Sliven)
=buresi Hanuš, 1923 Stara planina Mts.

ssp. zawadzkii Kraatz, 1854 E-Slovakia, NE-Hungary, W-Ukraine
=dissimilis Csiki, 1906 "Kom.Ung"(=W-Ukraine:Uzhgorod env.)
=seriatissimus Reitter, 1896 W-Ukraine:Carpathians:Hoverla Massif; ?"Marmaros"

ssp. ronayi Csiki, 1906 E-Slovakia("Kom.Zemplén & Ung"), NE-Hungary

ssp. simulator Kraatz, 1876 NE-Serbia:Majdanpek env.
=serbicus Hopffgarten, 1878 ?"Serbia"
=polychloros Beuthin, 1885
~tekijensis (Toševski, 1990); un NE-Serbia:Tekija:Djerdapska klisura, 50m(=Dunav right bank opposite to Orºova)



16.spasskianus Fischer, 1823 Altays Mts.to Transbaykalia; N-Mongolia

ssp. spasskianus Breuning, 1932 SC-Siberia:Krasnoyarsk env. to Transbaykalia; N-Mongolia; "Dauria"
=gryphus Motschulsky, 1846 "Baykal Lake"; "Sib.or.:Amaga"(LT)
=teicherti Mandl, 1963 N-Mongolia:"Dsun Charaa"

ssp. tungus Obydov, 1999 C-Siberia:Tunguska (Ilimpeya & Dovogpokan Riv.) & Podkam.Tunguska Riv.Basin

ssp. putus Motschulsky, 1846 C,W-Altays Mts.(Zmeinogorsk, Kolyvan; LT)

ssp. schestopalovi Plutenko, 1995 NE-Kazakhstan:Sarym-Sakty Mts.:Soldatovo env.

ssp. geblerianus Obydov, 1999 N-Altays Mts.:Barnaul env.(Barnaul)

ssp. saragaschensis Obydov, 1999 SC-Siberia:Hakassia:betw.Krasnoyarsk/Abakan:Saragash

ssp. cracens Obydov, 1999 E-Kazakhstan:Kalbinsky Mt.R.:Ognevka village

ssp. korostelevi Obydov, 2004 E-Kazakhstan:Ivanovsky Mt.R.(upper level of "Prokhodnoy Belok")

ssp. aeruginosiformis Breuning, 1932 West Sayans:Kazyr Suk River

ssp. microputus O.Berlov & Shilenkov, 1996 West & East Tannu-Ola Mt.R.:S-macroslope(Tuva:Khandagayty env.)

ssp. zinaidae Obydov, 1997 SE-Tuva:Sangilen Mts.:Kargy River valley:50:31N/97:01E, 1800m

ssp. shoriensis Obydov, 1999 SC-Siberia:Gornaya Shoriya(=SE of Novokuznetsk)



17.tarbagataicus Kraatz, 1878 E-Kazakhstan:Tarbagatay Mt.R., Saur Mts.; China:W-Xinjiang

ssp. tarbagataicus Kraatz, 1878 E-Kazakhstan:Saur Mts.("Tarbagatai"-probably inexact; LT)

ssp. urkasharensis Kabak, 2005 W-Xinjiang:Urkashar Mt.R.:near Kairakan Mt., 46:20:45N/84:06:45E; 1f

ssp. dshungaricus Csiki, 1927; nom.pro motschulskyi Lapouge E-Kazakhstan:Tarbagatay Mt.R.:C, E-part; "Dzhungaria" ~motschulskyi Lapouge, 1909; un "Dzhungaria" =fumidicolor Deuve, 1998 "Sibérie, Zlatoust"(=Zlatoust betw.Ufa/Chelyabinsk; by error); 1f =necincertus Deuve, 1998 "Sibérie orientale"(by error); 1m

ssp. pietrorattii Deuve, 1991 E-Kazakhstan:Tarbagatay Mt.R.:W-part(Urdzhar env.) =pseudogebleri Deuve, 1994 Tarbagatay Mts.:Alekseevka nr.Urdzhar



18.venustus Morawitz, 1862 Far East, N-Korea, NE-China

ssp. venustus Morawitz, 1862 Russia:Far East(E-Siberia:Bureya Mts.) =carinulatus Chaudoir, 1869; nec ? ?"Amur" =acutesculptus Chaudoir, 1877; nom.pro carinulatus Chaudoir ?"Amur" =christophi Kraatz, 1878 "Amur"

ssp. kaesongensis Imura, 1993 CW-Korea:Kaesong; 1f

ssp. hunjiangensis Deuve & Li, 2003 NE-China:Jilin:Hunjiang Xian:Linjiang:Mayihe



19.wulffiusi Morawitz, 1862 Far East, N-Korea, NE-China

ssp. wulffiusi Morawitz, 1862 Russia:Far East("Bai Possiet"=Pos’et SW of Vladivostok); NE-Korea:Chongjin, Hoeryong ~distinctus Haury, 1886, nec Rossi, 1792

ssp. dekraatzi Kraatz, 1881 NE-China:Liaoning(Liaodong env.), Jilin, Heilongjiang =jilinensis Deuve & Imura, 1990 NE-China:Jilin:Hunjiang Xian:Changbaishan Mts.:NW-slope:Bai-he =yonghongi (Li, 1992) Heilongjiang:Shangzhi:Mao’er Shan =yonghongianus Deuve, 1992; nom.pro yonghongi (Li) =pierremeyeri Müller, 2002 Heilongjiang:150km E Harbin:Zhangguangcai Mts., 45:10N/128:50E

ssp. opacipennis Reitter, 1895 NE-Korea:"Gensan"(=Wonsan) =flavifemoratus Imura & Yamaya, 1994 NW-Korea:Sinuiu env.:Mt.Chonma-san

ssp. odaesanus (Ishikawa & Kim, 1983) CE-Korea:Taebeaek-san Mt.R.:Mt.Odae-san

ssp. ssangnamensis (Kwon & Lee, 1984) NC-Korea:Huchang:Ssangnamdong; 1f

ssp. chonzeicus Deuve & Li, 2003 NC-Korea:Chagang-do:Huaren-ri:Mt.Chonze-san

ssp. wurhenensis Deuve & Li, 2003 NC-Korea:Chagang-do:Wurhen-ri:Mt.Sirubong-san

ssp. taebeagsanus (Ishikawa & Kim, 1983) CE-Korea:Taebeaek-san Mt.R.:Mt.Taebaek-san ~alpinus (Kwon & Lee, 1984), nec Paykull, 1790 Mt.Taebaek-san

ssp. liaoningensis Deuve & Imura, 1990 NE-China:E-Liaoning:Fengcheng:Fenghuang Shan; ?NW-Korea:Sinuiju:Chonma-san

ssp. onjinsanicus Rapuzzi, 1994 N-Korea:Hvanghe-pukto:Onjinsan Massif, 1000m(=ca.60km SE Pyongyang); 2m =myohyangicus Deuve & Li, 1999 NC-Korea:Hyangsan:Mt.Myohyang-san



20.zhubajie Imura, 1993 S-Shaanxi; SE-Gansu:Tianshui env.

ssp. zhubajie Imura, 1993 S-Shaanxi(Qinling Shan:Zhouzhi Xian);

ssp. caeliaquarius (Imura, 2004) SE-Gansu:Tianshui env.(Tianshui Shi, 1700m)

ssp. laojunensis Kleinfeld, 1999 W-Henan:Laojun Shan(Luangchuan, 33:43N/111:37E),Funiu Shan; SE-Shanxi:Zhongtiao Shan

ssp. shunwangping (Imura, 2006) S-Shanxi:Li Shan Mts.:SE slopes of Shun Wang Ping Mt., 35:24N/111:58E


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TRACHYCARABUS Géhin, 1885; type species Carabus scabriusculus Olivier
species: 9



1.besseri Fischer, 1822 NE-Romania, Moldavia("Süd-Podolien"), Ukraine
=dimorphus Lapouge, 1908
=pseudobesseri Lapouge, 1908



2.coriaceipennis Chaudoir, 1863 China:Shandong, Hebei, Shanxi, Shaanxi

ssp. coriaceipennis Chaudoir, 1863 CE-China:Shandong, Shanxi, Hebei; "Nordchina"

ssp. taibaimontanus Imura, 1994 C-Shaanxi:SW of Zhouzhi:Taibaishan Massif:S-slope:Houzhenzi



3.estreicheri Fischer, 1822 Ukraine("Proskurov"=Khmelnitskiy), SW-Russia; W-Siberia:Tyumen, Tobolsk
=adoxus Fischer, 1823 "Kaukasus"(error)
~modestus Fischer, 1823, nec Fabricius, 1801 "Dauria, Südrussland"
=jaegeri Ménétriés, 1827 ?Voronezh
=accuratus Chaudoir, 1848 "Bessarabia, Cherson"
~rufofemoratus Lomnicki, 1892, nec Motschulsky, 1865
~erythromerus Semenov, 1898, nec Dejean; nom.pro rufofemoratus Lomn.
~rubidofemoratus Csiki, 1927; un; nom.pro erythromerus Semenov
=basirufus Marcu, 1937



4.latreillei Fischer, 1822 C,E-Siberia, N-Mongolia, Far East, N-Korea; N-China:Shanxi to Liaoning

ssp. latreillei Fischer, 1822 C,E-Siberia(Irkutsk), N-Mongolia, Far East, NE-Korea; N-China:Shanxi, Nei Menggu, Hebei
=dejeani Fischer, 1822; nom.pro latreillei Fischer, 1822
=mannerheimi Dejean, 1826; nom.pro dejeani Fischer, 1822
~dimidiatus Fischer, 1844, nec Rossi, 1790 "Dauria"
~interruptus Fischer, 1844, nec Fabricius, 1790 "Dauria"
=sinensis Géhin, 1885 Chingan Mts.
=changaicus Lapouge, 1908 "Mts.on Siberia/Mongolia borders"
=inchanicus Lapouge, 1916 China:"Koukou-khoto"(=Nei Menggu:Honhot)
=tscheliensis Lapouge, 1916 NE-China:?"Tschili"(=Jilin)

ssp. mandschurensis Breuning, 1926 NE-China:"Mukden"(=Liaoning:Shenyang)

ssp. chonmanus Imura & Yamaya, 1994 NW-Korea:Pyongan-pukto:NE of Sinuiju:Mt.Chonma-san
=pyonganensis Deuve & Li, 1994 NW-Korea:N-Pyongan:NE of Sinuiju:Mt.Chonma-san

ssp. semicoriaceus Kraatz, 1881 NE-China:E-Liaoning:Liaodong Peninsula, Dandong env.; NW-Korea



5.mandibularis Fischer, 1827 Altays, Tarbagatay Mt.R., Saur Mt.R.; N-Mongolia; ?China:N-Xinjiang

ssp. mandibularis Fischer, 1827 Altays, Tarbagatay Mt.R., Saur Mt.R.; N-Mongolia; ?China:N-Xinjiang

ssp. buchtarmensis Kryzhanovskij, 1953 E-Kazakhstan:S-Altays:Bukhtarma River:Chernovaya



6.perrini Dejean, 1831 SE-Ukraine, Krym Peninsula, SW-Russia, Georgia

ssp. perrini Dejean, 1831 Krym Peninsula("Sudrussland, Krym")
=orbicollis Motschulsky, 1846 "Russ.mer."(LT)
=campicola Reitter, 1896 Krym

ssp. planus Géhin, 1885, nec Fabricius, 1792; nbm; nom.pro planatus Motschulsky SE-Ukraine, SW-Russia:Rostov, Maykop; Georgia; "S-Russia"
~campestris Fischer, 1822, nec Adams, 1817 "Tiflis"(=Tbilisi; ?inexact)
~parallelus Fischer, 1844, nec Faldermann, 1835 "S-Russia"
~planatus Motschulsky, 1850, nec Chaudoir, 1843; nom.pro parallelus Fisch. "S-Russia"
=transcaucasicus Csiki, 1927; nom.pro campestris Fischer "Tiflis"(=Tbilisi; ?inexact)



7.planarius Obydov, 1994 E-Kazakhstan:W-foot of Tarbagatay Mt.R.:20km S Taskesken:Sholpan



8.scabriusculus Olivier, 1795 SC-Europe, Carpathians, Balcans, Ukraine, S-Russia, W-Turkey

ssp. scabriusculus Olivier, 1795 Austria, Moravia, Slovakia, Hungary, W-Romania, Ukraine
=agrestis Creutzer, 1799 Moravia
~erythropus Fischer, 1822, nec Marsham, 1802
=hoffmanni Motschulsky, 1845

ssp. lippii Dejean, 1826 SE-Hungary; Romania, Moldavia
=?laevior Krynicki, 1830 "Moldavia, Bessarabia"
=curvatus Motschulsky, 1845 "Altays"(error)
~cruralis Lapouge, 1908; un
~costulatus Petri, 1912, nec Kraatz, 1889 "Transsylvania"
retyezaticus Csiki, 1927; nom.pro costulatus Petri "Transsylvania"
~macrolippii Lie, 1996; un C-Romania:S of Alba-Iulia:Sebeº-Alba env.

ssp. inapertus Motschulsky, 1850 Ukraine(Kharkov-LT), S-Russia
~longulus Lapouge, 1908; un

ssp. minutus Motschulsky, 1850, nec Fabricius, 1792; nbm W-Turkey:Bursa:Ulu Dag; "Georgia"(?error; LT)

ssp. bulgarus Lapouge, 1908 SC,SE-Romania, Bulgaria, Serbia, Bosnia, Macedonia, Eur.Turkey
~interruptus Born, 1902, nec Fabricius, 1775 S-Romania:Vlasca:Comana
=laceratus Csiki, 1927; nom.pro interruptus Born S-Romania:Vlasca:Comana



9.sibiricus Fischer, 1822 S,C-Siberia; Ukraine, Caucasus; ?NW-China:Xinjiang

ssp. sibiricus Fischer, 1822 SW,SC,SE,CE-Siberia; "Riddersk"(=Altays:Leninogorsk; LT)
=seriatoporus Fischer, 1827 Lena River

ssp. obliteratus Fischer, 1827; nom.pro obsoletus Fischer Altays, Sayans, Tarbagatay Mt.R.; N-Mongolia; "Siberia"
~obsoletus Fischer, 1823, nec Rossi, 1790 "Siberia"
=ledeburii Fischer, 1827 Altay
~tibialis Fischer, 1842, nec Paykull, 1798 "Ajagus"(=?Ayaguz 300 km S Semipalatinsk)
=riphaeus Motschulsky, 1846 Irtysh near Pianojarsk (=Semipalatinsk; LT)
=frater Kraatz, 1879 ?N-Mongolia
=frontosus Lapouge, 1908 "entre Khouldja/Ourga"

ssp. karelini Fischer, 1830 SW-Russia:Saratov, Orenburg, Samara; "Südrussland"

ssp. gvozdevae Obydov, 1997 Siberia:Hakassia:betw.Krasnoyarsk/Abakan:Saragash env.

ssp. kolosovi Zinoviev, 1997 S-Urals:Bashkiria:Mesyagutovo:Ay River bank; Perm:Kungur:Plekhanovo:Spasskaya Gora

ssp. bosphoranus Fischer, 1823 Taman Peninsula, Krym Peninsula, W-foot of Caucasus
~carbonarius Motschulsky, 1850, nec Rossi, 1790 S-Russia:?
=jailensis Breuning, 1932 Krym Peninsula:Jaila Mts.

ssp. tscherkessicus Korge, 1964 Caucasus; ?Armenia; ?NE-Turkey; ?LNF
~sibiriculus Mandl, 1955; un Caucasus:Kislovodsk, Teberda

ssp. errans Fischer, 1823 Ukraine; SW-Russia:Rostov; "Tauria"
=krynickii Fischer, 1827 Ukraine:Odessa
=cribricollis Motschulsky, 1846 "Sarepta"(=lower Volga basin; LT)
=maritimus Motschulsky, 1850 "Litt.m.Pont.:Odessa"
~acutangulus Eichler, 1924, nec Chaudoir, 1879 Rostov env.:Morskaya
~acutangulatus Csiki, 1927; un; nom.pro acutangulus Eichler Rostov env.:Morskaya
=retowskianus Mandl, 1955 SC-Ukraine:Odessa

ssp. haeres Fischer, 1823 N-Ukraine; SW-Russia:Voronezh, Tula, Ryazan’
~fossulatus Dejean, 1826, nec Quensel, 1806

ssp. rybinskii Reitter, 1886 W-Ukraine:Ternopol

ssp. pseudobliteratus Korell & Kleinfeld, 1982 NW-China:Xinjiang:"Ake-Tei-A-Zhu-Pai Temple"(=?nr.Korla)


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MIMOCARABUS Géhin, 1885; type species Carabus maurus (Adams)
species: 5



1.elbursensis Breuning, 1946 N-Iran:Elborz Mts.:?"Mt.Sulaiman"(=Taht-i Suleiman)



2.khorasanensis Deuve, 1993 NE-Iran:"Ala-Dagh, Budschnurd"(=Bojnurd); 30km N Ghuchan:37:23N/58:31E



3.maurus (Adams, 1817) Armenia, Georgia; C,E-Turkey; Cyprus; N,C-Iran; Syria, Lebanon

ssp. maurus (Adams, 1817) NE-Turkey:Kars; Armenia; Georgia("Tiflis"=Tbilisi)
=gemellus Fischer, 1844 "Caucasus"
=discoideus Reitter, 1880
=alagoensis Kraatz, 1896 "Mts.S of Araxes valley:Alagoes"(=Mt.Aragats Massif)
=maurulus Mandl, 1955 Caucasus

ssp. paphius Redtenbacher, 1843 Cyprus, SE-Turkey, N-Iran, N-Iraq, Syria, Lebanon

ssp. osculatii Osculati, 1844 W-Iran southwards to Shiraz env.
=schirasensis Breuning, 1968 CW-Iran:S of Shiraz

ssp. hochhuthi Chaudoir, 1846 C,E-Turkey:Amasya, Konya, Eskisehir, Gümüshane; "Armenia"(?NE-Turkey:Bayburt env.)
=geminatulus Reitter, 1896 Amasya

ssp. calosomoides Reitter, 1896 S-Turkmenistan:Kopet Dag Mts.; NE-Iran; "Transcaspia:Hadscha-Kala"(=?)
=demetrii Semenov, 1903 N-Iran:Mazenderan("Mazanderan:Sefidab River")
=caspicola Rapuzzi, 2005 N-Iran:Mazandaran:E of Kelardasht, 36:30N/51:15E
ssp. hermonensis Schweiger, 1970 Antilibanon Mts.:Hermon Massif



4.pumilio Küster, 1846 Armenia, Caucasus Minor; NE-Turkey:Ararat Massif
=araratensis Kraatz, 1896 Mt.Ararat



5.roseni Reitter, 1897 NE-Iran; Turkmenistan:KopetDag Mts.

ssp. roseni Reitter, 1897 NE-Iran:"Haidanabad"(=?Torbat-e Heydariyeh; LT); Turkmenistan:KopetDag Mts.

ssp. hemicalosoma Semenov, 1903 NE-Iran:Gorgan env., Bojnurd env.("Budshnurd Mts.)

ssp. gonbadensis Deuve, 1991 NE-Iran:NE of Gorgan:Gonbad-e Kavus; 70km NE Gonbad-e-Kavus:37:26N/55:23E


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RHIPOCARABUS Reitter, 1896; type species Carabus alysidotus Illiger
species: 1



1.alysidotus Illiger, 1798 NW,CW-Italy; Elba Island; SE-France:Alpes-Maritimes to Hérault; "Italy"
=pomptinus Lapouge, 1916 Italy:Maremma
=stagnalisaequalis Lapouge, 1916 Hérault:Lattes nr.Montpellier
=stagnalis Breuning, 1932; nom.pro stagnalisaequalis Lapouge
=bucheti (Ochs, 1949)


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ARCHICARABUS Seidlitz, 1887; type species Carabus nemoralis Müller
species: 9



1.gotschi Chaudoir, 1846 Turkey, Armenia, W-Syria

ssp. gotschi Chaudoir, 1846 Armenia, NE-Turkey:Kars, Erzurum; "Armenia"
=maljushenkoanus Breuning, 1926 Armenia:Yerevan, Sevan Lake; "Göktschai"(=Sevan)
=horioni Korge, 1968 Kars:betw.Susuz/Haskoy:Kisir Dagi
=naumanni Breuning, 1971 Kars:"Merdenik"(=Göle)

ssp. congruus Motschulsky, 1865 C-Turkey:Amasya, Samsun, Nevºehir, Ankara, Cankiri, Tokat, Sivas, Yozgat, Kayseri
~turcicus Motschulsky, 1865, nec Fabricius, 1792 LNF
=ottomanus Reitter, 1896 Amasya
=diversus Schweiger, 1969 Nevsehir:Avanos
=ilgazicus Breuning & Ruspoli, 1970 Cankiri:betw.Ilgaz/Cankiri:15km S Ilgaz

ssp. pseudogotschi Breuning, 1926 C-Turkey:Konya(Sultan Dag:nr.Aksehir), Ankara, Eskisehir
=inonunus Cavazzuti, 2003 Eskisehir:Inönü, Sögüt env.
~kochisarensis Lassalle, 2003; un Ankara:Sereflikochisar

ssp. regismontium Schweiger, 1964 NC-Turkey:Kastamonu:Ilgaz Dag:"Juniperus zone"
=subnivalis Schweiger, 1964 Kastamonu:Ilgaz Dag:"alpine zone"

ssp. pseudorugosus Machard, 1994 E-Turkey: Bitlis(Resadiye), Artvin, Ezzurum, Gümüshane, Sivas, Bingöl
=nicolae Auvray, 1995 Artvin:Kilickaya
~koyulicus Lassalle, 2003; un Sivas:Koyulhisar

ssp. caramanus Fairmaire, 1886 SE-Turkey:Antakya("Akbes"=E of Hassa); W-Syria
=mollis Lapouge, 1924 Antakya:Akbes
=depressipennis Mandl, 1955 LNF(unlabelled HT)

ssp. unctus Lapouge, 1908 SC-Turkey:Adana, Konya, Tarsus, Kahramanmaraº, Kayseri; "Taurus"
=bulghardaghensis Breuning, 1926 SC-Turkey:"Burna, Bulghar Dagh"(=Bolkar Dag)
~goeksunensis Breuning, 1966; un Kahramanmaras:Göksun env.; Kayseri:Erciyas Dag
=eregliensis Breuning, 1974 Konya:Eregli
=demelti Heinz, 1977 Mersin:Namrun:Camliyayla; Adana
=alayliensis Cavazzuti, 2006 Adana/Kayseri:betw.Dogabeyli/Bakirdagi:Kücükgezbeli



2.monticola Dejean, 1826 SE-France & NW-Italy:Alps

ssp. monticola Dejean, 1826 SE-France:Isére, Alp.Hte.Prov.("Basses Alpes"), Alp.Marit.; NW-Italy:Liguria
=asperus Beuthin, 1892 ?"Basses Alpes"(=Alpes-Hte-Provence)
=jordani Beuthin, 1892 ?"Basses Alpes"(=Alpes-Hte-Provence)
=maritimensis Born, 1923 Alpes Maritimes
=megasomus (Tarrier, 1972) Alpes Maritimes:?lower Vésubie valley
~bleonensis Raynaud, 1973; un Alpes-Hte-Provence:Bléone valley:Prads

ssp. liguricus Born, 1898 NW-Italy:Alpi Liguri(Colle dei Signori)

ssp. fontanae Born, 1906 NW-Italy:Monte Generoso, Biella env.
=biellensis Born, 1923 Biella env.

ssp. roccae Born, 1923 NW-Italy:Monte Viso, Albergian, Val di Susa, Col Coupé; betw.Savona/Torino
~sturanii Mandl, 1966; un Liguria:Savona env.:Monte San Giorgio



3.montivagus Palliardi, 1825 Balcans, Romania, Hungary, SE-Slovakia

ssp. montivagus Palliardi, 1825 C,W,E-Bulgaria; C,W-Romania("Banat"), E-Serbia, E-Macedonia; NE-Greece, Eur.Turkey
=kalofirensis Apfelbeck, 1904 Bulgaria:"Kalofer"(=Kalofer Planina Mts.).
~ponticus Apfelbeck, 1904, nec Deyrolle, 1869 E-Bulgaria:Burgas
~rosalitanus Apfelbeck, 1904; un "Balkan Mts.:Rosalita Pass env.(=Stara Planina Mt.R.)
=bulgaricus Csiki, 1927; nom.pro ponticus Apfelbeck E-Bulgaria:Burgas

ssp. vellepiticus Hampe, 1850 Croatia(Velebit Mts.), Bosnia, Montenegro; N-Albania
~illyricus Kraatz, 1880, nec Kraatz, 1878 ?"Ternowaner Wald"(=Slovenia:Trnovski Gozd)
=illyriensis Géhin, 1885; nom.pro illyricus Kraatz ?"Ternowaner Wald"(=Slovenia:Trnovski Gozd)
=sutomorensis Reitter, 1885 Montenegro:"Sutomore"
=leonhardi Born, 1904 ?"N-Hercegovina"
=medius Lapouge, 1908 "Romania, Bulgaria, Serbia, Bosnia & Croatia"
=goljensis Born, 1910 W-Bosnia:Golja Planina
=boszdaghensis Breuning, 1966 W-Turkey:"Bos Dagh"(=Boz Dag; error)

ssp. blandus Frivaldsky, 1865 S-Slovakia, Hungary("Kom.Abauj-Torna & Heves")



4.nemoralis Müller, 1764 Europe:S-Spain - British Isles – Scandinavia - Eur.Russia; introduced:N.America, Kazakhstan

ssp. nemoralis Müller, 1764 Europe:France - Brit.Isles – Scandinavia - Eur.Russia; Germany; introd.:N.America, Kazakhstan
=foetens Voet, 1778
~nigrescens Letzner, 1850; un "Silesia"
~virescens Letzner, 1850; un "Silesia"
~tristis Dalla Torre, 1877; un Oberösterreich
~krasae Roubal, 1903; un SW-Bohemia:"Klattau"(=Klatovy)
~brunnipes Lapouge, 1908; un
~lucidus Lapouge, 1908, nec Duftschmid, 1812 ?N,C-France
~canadensis Lapouge, 1908, nec Melsheimer, 1853 N.Brunswick & Newfoundland(introduced)
~deletus Lapouge, 1908; un "Russia & Hungary"
~auratus Heuer, 1926, nec Linné, 1758 Frankfurt a.M.
~lestagei Basilewsky, 1930; un
=nicollei Bourgin, 1947 Aube:?Ft.du Grand Orient
=nigrotinctus Mandl, 1955
~pseudomontivagus Mandl, 1961; un "Kärnten:Weidisch:Javornik"
=verdonensis Puisségur, 1961 Alpes-Hte-Provence/Var:Verdon valley:Les Dourbes
=colasi (Bourgin, 1963) Massif des Maures
=miolansicus Tarrier, 1965 Esteron valley
=cantalicus Jeanne, 1970 Cantal:Plomb du Cantal, 1850m
~aubersoni (Raynaud, 1973); un Switzerland:Pomy
=borealensis (Raynaud, 1973) England:Oundle, Ashton Wold
~cantalicus Machard, 1974, nec Jeanne, 1970 C-France:Cantal:?
=fayardensis Machard, 1974 Alpes-Hte-Provence:Lardier
~freneyi Tarrier, 1975; un Maine-et-Loire:Bois du Mouliherne
~montisdiniensis Tarrier, 1975; un Alpes-Hte-Provence:Les Dourbes
=waltersachi Mandl, 1984 Austria:Steiermark:Rotemanner Tauern:Strechengraben

ssp. prasinotinctus Heyden, 1880 SW-France, NC-Spain; Lozére & N-Spain
=contractus Géhin, 1885 Hautes-Pyrenées
=kraatzianus Beuthin, 1889 ?"S-France"
=nisseni Beuthin, 1889 ?"S-France"
~pulcherrimus Beuthin, 1889; un ?"S-France"
=atavus Lapouge, 1908 LNF(unlabelled HT)
~discolor Lapouge, 1908; un
~meridionalis Lapouge, 1908, nec Born, 1905 Ariége
=pascuorum Lapouge, 1908 S-France:Cévennes; Lozére:St.Germain-de-Calberte(NT)
=quinqueseriatus Lapouge, 1908 Landes
~setosus Lapouge, 1910; un
~meridianus Csiki, 1927, nec Linn., 1767; nom.pro meridionalis Lapouge Ariége
~venustus (Raynaud, 1973), nec Morawitz, 1862 Lot:Leymes
~litigiosus Tarrier, 1975; un Aude:Ft.de Monthaut nr.Rennes-les-Bains
?manae Devecis, 1996; un Corréze:Monédi?res Massif

ssp. lamadridae Born, 1895 NW-Spain:Lugo, Oviedo, Leon, Coruna; "Asturia:Llanos"
~indigotinctus Codina, 1927; un N-Spain:Burgos:Pancorbo
~layrei Tarrier, 1975; un NW-Spain:Lugo:Puertomarin



5.phoenix Lapouge, 1924 SW-Lebanon:Saida, Djebel Ech Cheik Mts.; ?SW-Syria



6.pseudomonticola Lapouge, 1908 S-France:Pyrenees-Orientales(Serralongue; NT); NE-Spain:Gerona, Barcelona
=sorredensis Reymond, 1935 Pyrenees-Orientales:Foret de Sorréde
~catenus (Raynaud, 1973); un "Casteil"
~lunatus (Raynaud, 1973); un "Col des Ares"
~hispanicus Tarrier, 1975; un Barcelona:Montserrat
=pseudopascuorum Tarrier, 1975 Barcelona:N.S.de Queralt nr.Berga



7.rossii Dejean, 1826 Italy:Piemonte to Calabria
=dragonettii Rozzi, 1839 Gran Sasso Massif
~castaneipennis Costa, 1857, nec Ménétriés, 1832
~minor Pirazzoli, 1871, nec Kraatz, 1860 Abruzzi Mts.
=pirazzolii Géhin, 1876; nom.pro minor Pirazzoli Abruzzi Mts.
=costae Géhin, 1885; nom.pro castaneipennis Costa
=stoecklini Lopez, 1891 Livorno
~intermedius Porta, 1923, nec Dejean, 1826
=portae Csiki, 1927; nom.pro intermedius Porta



8.steuartii Deyrolle, 1852 NW-Spain:Orense, Zamora, Lugo, Léon; N-Portugal:Braga to Guarda("Oporto env."=Porto)
=heydeni Oliveira, 1876 Portugal:?Serra de Gerez
=barrosi Lapouge, 1908
=fusarius Lapouge, 1908
=moreli Lapouge, 1908 Spain
=nodicornis Lapouge, 1908
=fabriciiruspolii Breuning, 1966 NW-Spain:Léon:Pena Rubia
=durani Raynaud, 1974 NW-Spain:Lugo:Samos
=bastiani Lassalle, 1982 Portugal:Guarda:Manteigas env.



9.victor Fischer, 1836 SE-Bulgaria, NW, NC, NE-Turkey, SW-Georgia, ?W-Armenia

ssp. victor Fischer, 1836 NC, NE-Turkey:Ordu to Artvin, SW-Georgia(Akhaltsikhe)
=motschulskyi Kolenati, 1845 "Tiflis"(=Tbilisi)
=inconspicuus Chaudoir, 1846 Trabzon, Erzurum, Artvin, Kars; "Armenia"(=NE-Turkey)
=falsarius Reitter, 1896 Armenia:"Alagoes"(=Aragats; ?by error)
=nanus Kraatz, 1896
~obtritus Lapouge, 1924, nec Chaudoir, 1857 "Urals"(by error)
=transeuns Lapouge, 1924 NC, NE-Turkey:Ordu, Sivas; "Bosporus, Skutari"(by error)
=dissolutus Csiki, 1927; nom.pro obtritus Lapouge "Urals"(by error)
~akkusanus Breuning, 1965; un Ordu:Akkus
~cordithorax Blumenthal & Breuning, 1967; un Rize:Ayder nr.Camlihemsin
=montanellus Heinz, 1967 Artvin:Yalnizçam Pass
=loebli Schweiger, 1969 Trabzon:Gesirmendereº valley
=pseudotransenus Machard, 1979 Ordu:betw.Gölköy/Mesudiye
=hortulanus Darge, 1994 Rize:Ovit Pass
=subminimus Machard, 1995 Trabzon:S of Caykara:Soganli Pass, 2300m
=borckanus Cavazzuti, 2003 Artvin:Borcka
~torulensis Lassalle, 2003; un Gümüshane:W of Torul

ssp. wiedemanni Ménétriés, 1836 NW, NC-Turkey:Istanbul to Sinop; SE-Bulgaria; NE-Greece:Alexandroupoli; "Turkey"
=vaitoiani Strasser, 1900 "Phillipoppel"(=Plovdiv)
=burgassiensis Apfelbeck, 1904 SE-Bulgaria:Burgas env.
=bythinus Lapouge, 1908 "Anatolia:Kadikoy"(=?Bospor region)
=anceps Lapouge, 1924 LNF(?Amasya/Samsun region)
=winklerianus Breuning, 1933 Bursa:Ulu Dag Massif
=paphlagoniensis Breuning, 1954 Kastamonu
=goekdaghensis Mandl, 1955 Kocaeli:Goek Dag (=S of Izmit)
=cekirgesis Schweiger, 1964 Bursa:Havuzlu park nr.Cekirge
~nilufer Schweiger, 1964; un Bursa:Havuzlu park nr.Cekirge
~patruelis Schweiger, 1964; un Kastamonu:Ilgaz Dag:grasslands
=besucheti Schweiger, 1965 Bolu:pass betw.Bolu/Kaynasli
~mussardianus Breuning, 1965; un Sinop:Dranaz Dag
~akcakocaensis Blumenthal & Breuning, 1967; un Bolu:Akcakoca env.:Kaplandere Dagi
=mengenicola Schweiger, 1969 Bolu:SW of Mengen
~zindangensis Lassalle, 2003; un Sinop:Zingan Dag, Erfelek, Yenikonak, Kastanelik


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ISCHNOCARABUS Kraatz, 1877; type species Carabus tenuitarsis Kraatz
species: 2



1.cychropalpus Peyron, 1858 S-Turkey:Adana("Cydnus River bank"=?Ceyhan River), Antakya
=cychropalpulus Mandl, 1955 Adana



2.tenuitarsis Kraatz, 1877 C-Turkey

ssp. tenuitarsis Kraatz, 1877 C-Turkey:Amasya, Tokat, Çorum
=cottarellii Breuning & Ruspoli, 1970 "betw.Amasya/Çorum:Mongitosu"(=Çorum:Mecitözü)
=balgoezensis Deuve & Charet, 2004 C-Turkey:Çorum:Balgöze

ssp. ankarensis Breuning, 1961 C-Turkey:road Ankara/Çankiri:20km N Ankara
~cankiriensis Breuning, 1962; un Çankiri:betw.Çankiri/Ilgaz
=prasinosomatus Rapuzzi, 1991 Çankiri:Çankiri env.

ssp. auvrayi Charet, 1992 Kirºehir:Çiçekdagi pass E of Ankara
=rubizzaniorum Cavazzuti, 1994 C-Turkey:Kirºehir:Mucur


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LIMNOCARABUS Géhin, 1876; type species Carabus clathratus Linné
species: 1



1.clathratus Linné, 1761 N,C-Europe, Brit.Isles, Balcans, Turkey, Russia, Central Asia, Siberia, Far East, Japan

ssp. clathratus Linné, 1761 N,C-Europe, British Isles, Russia, Central Asia, Siberia; LNF(?N-Germany)
=aeneopunctatus DeGeer, 1774
=longus Voet, 1778
=adspersus Gmelin, 1788
~eversmanni Motschulsky, 1846, nec Fischer, 1832 "Ichym"
~detritus Letzner, 1850; un "Silesia"
~cupreus Schilsky, 1888, nec Linné, 1758 N-Germany:Borkum Island, Warnemünde, Zingst
=jansoni Kraatz, 1890 Scottland
~multipunctatus Kraatz, 1890, nec Linneaus, 1758 "Westpreussen"
~borealis Born, 1908, nec Paykull, 1790
=foveolatoseriatus Reitter, 1896 Altays
=boreus Csiki, 1927; nom.pro borealis Born
~aeratus Culot, 1986; un

ssp. stygius Ganglbauer, 1890 Azerbaydzhan:Lenkoran env.; N-Iran

ssp. auraniensis Müller, 1902 Austria, S-Moravia, Slovakia, Hung., SE-France; Italy; Balcans("Fiume"=Rijeka); Cauc., Armen., Turkey
~laccophilus Reitter, 1896; un "Caucasus:Mahmutli"
=arelatensis Lapouge, 1903 SE-France:Camargue, Montpellier
=antonellii Luigioni, 1921 C-Italy:Latium:Maccarese,Fiumicino, Roma env.
=epirensis Purkynì, 1928 NW-Greece:Ipeíros:"Janina"(=Ioánnina)

ssp. maacki Morawitz, 1862 Far East(Hanka Lake):Ussuri, Sikhote-Alin; Korea; Japan:N,C-Honshu
=aquatilis Bates, 1883 Japan:Honshu:Shimonosawa Lake(=?Nagano:Suwa-ko Lake)
=viridipunctus Kraatz, 1886 Far East:Suyfun River(=W of Vladivostok)


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TACHYPUS Weber, 1801; type species Carabus auratus Linné
species: 3



1.auratus Linné, 1761 W,NC-Europe

ssp. auratus Linné, 1761 W,NC-Europe:NC-Spain:Viella; SC,C,N-France to Bohemia, ?W-Poland; "Sweden"(error)
=sulcatus DeGeer, 1774
~coeruleomicans Letzner, 1850; un
~contortus Letzner, 1850; un
~picipes Letzner, 1850; un
=laniarius Gistl, 1857
~nigripes Dalla Torre, 1877; un
~opacoviridis Dalla Torre, 1877; un
~viridiauratus Dalla Torre, 1877; un
=brullei Géhin, 1885
=catalaunicus Géhin, 1885 "Champagne"
=obscuricornis Beuthin, 1894
=worleei Beuthin, 1894 "Harz"
~rotundatus Born, 1895, nec Paykull, 1798 Col de Balme nr.Chamonix
=auratoides Reitter, 1896 "Vogesen"(=Vosges Mts.)
=perauratus Reitter, 1896 Austria:Vorarlberg:Dornbirn
=sulcatissimus Lapouge, 1898
=confluentinus Bockleth, 1904 C-Germany:Rheinland:Koblenz
=laticollaris Bockleth, 1904
=quadricostatus Bockleth, 1904
=labittei Clément, 1904 Paris env.
~perviridis Born, 1915; un; error for perauratus Reitter
~anthracinus Barthe, 1921; un "Pyrenees"
=bettingeri Barthe, 1921 Savoie:St.Pierre d’Albigny
=brunieri Barthe, 1921 Aude:Salvezines
~erythropus Barthe, 1921; un "C-Pyrenees"
~dufouri Barthe, 1922, nec Dejean, 1829 Auvergne:Clermont-Ferrand
~barthei Lebis, 1924; un
=navarricus Lapouge, 1924 ?"W-Pyrenees"; ?"Navarra"
=hilairei Gavoy, 1925 Pyrenees:Val d’Aran
=dufouranus Csiki, 1927; nom.pro dufouri Barthe Auvergne:Clermont-Ferrand
~nigricatus Csiki, 1927; un; nom.pro anthracinus Barthe "Pyrenees"
~rutilipes Csiki, 1927; un; nom.pro erythropus Barthe "C-Pyrenees"
=clementi Sirguey, 1931
~clermonti Sirguey, 1931; un
~meyeri Sirguey, 1931, nec Born, 1898
=rivalieri Bourgin, 1948 Seine-et-Marne:?
=concyri Machard, 1973
~plonnieri Machard, 1973; un
=isignyensis Machard, 1977
~margaritae Hecq, 1996, un Marne/Meuse:Ft. d’Argonne(Chatrices S of Sainte Menehould - TNF)

ssp. honnoratii Dejean, 1826 SE-France:Alpes-de-Haute-Provence(?"Mts.of Basses Alpes:Banon"), Vaucluse, Var
~atripes Géhin, 1885; un "Basses Alpes"(=Alpes-de-Haute-Provence)
~nicolasi Chobaut, 1897, nec Reitter, 1888 Vaucluse:Mt.Ventoux
=ventouxensis Nicolas, 1904 Vaucluse:Mt.Ventoux
=hnicolasi Chobaut, 1908; ?nom.pro nicolasi Chobaut Vaucluse:Mt.Ventoux
~baeri Bleuse, 1914, nec Ménétriés, 1851 "Basses Alpes"(=Alpes-de-Haute-Provence)
=diversicolor Bleuse, 1914 "Basses Alpes"(=Alpes-de-Haute-Provence)
~olivarius Nicolas, 1919; un Alpes-de-Haute-Provence:Digne
~nicolasianus Csiki, 1927; un; nom.pro nicolasi Chobaut Vaucluse:Mt.Ventoux
=fabrei Colas, 1962 Vaucluse:Mt.Ventoux
=vadoni Colas, 1962 Var:Rians
=aufrerei Colas, 1964 Alpes-de-Haute-Provence:Les Dourbes
~diensis Tarrier, 1975; un Drôme:Foret de Saou
~blayeulensis Machard, 1977; un Alpes-de-Haute-Provence:Mt.de Blayeul
~lardierensis Machard, 1977; un
~lurensis Machard, 1977; un
~grimonensis Machard, 1977; un
~malcorensis Machard, 1977; un
~seirinensis Machard, 1977; un
~ignescens Darnaud, 1980; un Vaucluse:Mt.Serein
~astridae Fiévet, 1991; un
~camardi Darnaud, 1995; un Vaucluse:Buoux

ssp. lotharingus Dejean, 1826 SC,SE-France:Gard, Herault, Aude, Vaucluse, Bouch.du Rh., Ardeche; Moselle:Metz(error)
=siculus Ragusa, 1881 "Sicilia"(error)
=monspessulantus Lapouge, 1898 Gard:Montpellier
=transiens Cleu, 1952
=incognitus Bourgin, 1953 Hérault:?
=puisseguri Bourgin, 1953 Hérault:Les Thiéres
=gangensis Schaefer, 1962 Hérault:Ganges
=billieriensis (Schaefer, 1969) Hérault:Les Billieres
=virenquensis (Schaefer, 1969) Aveyron:La Virenque
=avenionensis (Schaefer, 1973) Vaucluse:Avignon
=larzacensis (Schaefer, 1973) Hérault:Le Caylar
=mourguesi (Schaefer, 1973) ?Hérault:Cros env.:La Virenque
=sambucensis (Schaefer, 1973) Bouches-du-Rhone:Le Sambuc
?gagates Forel & Leplat, 1995 Hérault:Brassac

ssp. lasserrei Doué, 1855 SW,SC-France(Dep.Ardeche)
=ligericinus Fairmaire, 1886 Lozére:S-part
=sudresi Lapouge, 1898 Lozére:Mt.de la Margeride
~ruthenus Lapouge, 1908; un
=magdelainei Bleuse, 1914 Ardeche
=ispaniacus Nicolas, 1919 Lozére:Ispagnac
=pseudolotharingus Barthe, 1921 Aude:?
~taranicus Cleu, 1952; un Ardeche
~ligericinulus Mandl, 1955; un
=carouxensis (Schaefer, 1969) Hérault:Mt.Caroux
=millavensis (Schaefer, 1969) Aveyron:?
~mouriesi (Schaefer, 1969); un
~rabili Raynaud, 1971; un Tarn:Foret de Gresigne
=mazametensis Prunier, 1999 Tarn:S of Mazamet:3km of Y?s
?agoutensis Bidault & Camard, 2003; un Hérault:upper Agout valley:"Cambon et Salvergues"



2.cancellatus Illiger, 1798 W,N,C-Europe, Balcans, S-Scandinavia, W-Russia, Siberia

ssp. cancellatus Illiger, 1798 N-Germany, S-Scandinavia, Poland, N-Moravia, Ukraine, W-Russia, Siberia; ?"Ost-Preussen"
=tuberculatus Dejean, 1826 "N-Germany"
=conjunctus Lomnicki, 1892
=sajanensis Reitter, 1896 "Ostsajan"(=East Sayans)
~anderseni Beuthin, 1896; un
~bicolor Beuthin, 1896, nec Walker, 1866
=conspersus Lapouge, 1902 ?"Russia"; ?Byelorussia:Baranovichi
~rossicus Lapouge, 1902; un C-Russia:?
~adelphus Kolbe, 1912, nec Rost, 1892 "Mark Brandenburg"
=amitinus Kolbe, 1912
=avunculus Kolbe, 1912 "Westpommern:?Negast nr.Stralsund"
~fallax Kolbe, 1912, nec Olivier, 1795 CW-Poland:"Krossen"(=Krosno)
=genuinus Kolbe 1912
=marchicus Kolbe, 1912 Berlin env.
=rauterbergi Kolbe, 1912 Niedersachsen:Hildesheim
=saxonicus Kolbe, 1912 "Sachsen:?"
=kniehopfi Langenhan, 1913 "Pommern:Stolp"
~balticus Bernau, 1915; un Latvia:"Lennewaden nr.Riga"
~charcoviensis Bernau, 1915; un Ukraine:Kharkov
=pratensis Bernau, 1915 "Südrussland"(error)
=suspicax Everts, 1915 Holland:?
~rapax Kolbe, 1915; un N-Moravia:Paskov
=wankae Sokolar, 1919 "Teschen & Saybusch"
=munsteri Born, 1926 SE-Norway:?
=dolosus Csiki, 1927; nom.pro fallax Kolbe CW-Poland:"Krossen"(=Krosno)
~nowotnyi Eidam, 1930; un "Emanuelsegen"
~semnonicus Csiki, 1927; un; nom.pro adelphus Kolbe "Mark Brandenburg"
=kuntzeni Eidam, 1930 Denmark:"Jutland:Sandbl.Leijord, Sandet, Muldjord, Klitplantage, Svingelbjerg"
=obscuriusculus Eidam, 1930 "Oberschlesisches Industriegebiet"
=ingulensis Eidam, 1941 Ukraine:"Elisabethgrad, Cherson, Charkow"

ssp. scythicus Schaum, 1857 Romania & W-Ukraine:N,C-Carpathians; ?Carpathians of C-Romania
=bucsecsianus Born, 1902 "Gipfelregion des Bucsecs"(=Bucegi Mt.R.; alpine form)
~pseudoscythicus Lapouge, 1902; un
~pseudotuberculatus Lapouge, 1902; un ?Carpathians of NC Romania
~transsylvanicus Petri, 1912; un "Transsylvania:Schulergebirge, Schässburg"(=Sighiºoara)
=assiduus Kolbe, 1913 Fagaraº Mt.R.:Mt.Negoi
=bucovinianus Kolbe, 1913 "Bucowina"(=NE-Romania)
~gracilis Kolbe, 1913, nec Küster, 1846 "Galicia:Radymno"
~?infernalis Kolbe, 1913; un "Ungarn"(=?W-Romania)
=moldavicus Kolbe, 1913 Bacau
=oligoscythus Kolbe, 1913 NE-Romania:"Berlad valley"(=Birlad River)
=oriundus Kolbe, 1913 "Czernowitz & Bojan"(=W-Ukraine:Chernovtsy)
=romaniacus Kolbe, 1913 "Romania:Parban"
~dobrudschensis Bernau, 1913; un Dobrogea Lowland(?error)
~subfallax Bernau, 1913; un "Bukowina:Bojan"
~scythicoides Bernau, 1913; un "Walachei"
~vorax Bernau, 1913; un "Kronstadt"(=Braºov)
~subscythicus Lie, 1996; un C-Romania:NE of Simeria:Sibot env.

ssp. durus Reitter, 1896 W-Carpathians:Slovakia("Trencsener Bergen"=Mts.nr.Trenèín), SW-Ukraine, NC-Romania
=pseudograniger Reitter, 1896 "Trencsén"(=SW-Slovakia:Trenèín)
=ungensis Csiki, 1905 ?"Kom.Ung"(=W-Ukraine:Uzhgorod env.)
~carpathicus Kolbe, 1913, nec Palliardi, 1825 "Szinever & Nadvorna"(=?Carpathians of W-Ukraine)
=enitens Kolbe, 1913 "Galicia"
=strictus Kolbe, 1913 "Galicia & Karpathen"

ssp. sulinensis Born, 1902; ?non priority NE-Romania("Letea nr.Sulina/Dunarea confluence"), Moldavia, Krym Peninsula
=?semistriatus Fischer, 1823; priority "Caucasus"(error)
=scabridus Eidam, 1941 LNF(?Moskva, Ryazan’, Kazan’)

ssp. emarginatus Duftschmid, 1812, nec Olivier, 1795; nbm NE,NC-Italy, W-Slovenia, SW-Switzerland, SE-Austria, SE-Croatia, NW-Bosnia; ?"Krain"
~affinis Duftschmid, 1812, nec Schrank, 1781
~oblongus Sturm, 1815, nec Fabricius, 1792 LNF(?Italy/Slovenia/Austria borders region)
=dahli Heer, 1841 Alpi Orobie:Mt.Generoso
=duftschmidi Géhin, 1876; nom.pro affinis Duftschmid
=trentinus Kraatz, 1877 "Trient"(=Trento)
=bohatschi Reitter, 1881 Lago Maggiore
~tridentinus Bertolini, 1887; ?iss of trentinus Kraatz
=penninus Lapouge, 1896 "Monte Rosa"
~sequensi Beuthin, 1898; un CW-Croatia:Gospic
~generoso Born, 1900; ?nud Alpi Orobie:Mt.Generoso(alpine form)
~collaris Lapouge, 1902; un "S-Tyrolia:?
=generosensis Born, 1906 Alpi Orobie:Mt.Generoso(alpine form)
=aurosplendens Born, 1910 ?SE-Croatia, ?NW-Bosnia
~karstianus Bernau, 1911; un SW-Slovenia("Heidenschaft"=Ajdovšèina)
~tolminensis Bernau, 1911; un Italy/Slovenia:Isonzo valley:Tolmin env.
=luganensis Born, 1914 "Ticino:Locarno & Lugano"
=picciolii Bernau, 1915 "N-Appeninnes"
=ticinus Born, 1920 "Tessin"
=poschiavinus Born, 1922 Lombardia:Bernina Massif:Val Poschiavo
~obscuripennis Mandl, 1961; un "Kärnten:Waidischtal:Hudajama-Graben"
~planitialis Ghiretti, 1994; ?un Lombardia:Mantova

ssp. carinatus Charpentier, 1825, nec Duftschmid, 1812; nbm France, NW,NC-Spain, W-Switzerland("Genf"=Genéve), W-Germany, Belgium, S-Holland
=fusus Palliardi, 1825 Savoie
=dolens Kraatz, 1879 "Pyrenees"
=tarnensis Géhin, 1885 Tarn
=aveyronensis Beuthin, 1895 Aveyron
~nigrinus Beuthin, 1896, nec Motschulsky, 1865
~pseudocarinatus Beuthin, 1896; un "Rheinland"
=astur Lapouge, 1898 "Asturia"
=celticus Lapouge, 1898 "Intérieur de France"
=crassus Lapouge, 1898 "Batz & Le Croisic"
=misellus Lapouge, 1898 "Margeride Mts."; alpine form
=pyrenaicus Lapouge, 1898 S-France:?
~acicularis Lapouge, 1902; un"S-France"
~asidoides Lapouge, 1902; un "S-France"
~inornatus Lapouge, 1902; un
~confinis Lapouge in Barthe, 1909; un
=progressivus Kolbe, 1912 "Westfalen"
~subcarinatus Kolbe, 1912; un "Rheinprovinz"
=houlberti Bleuse, 1913 Ille-et-Vilaine
=nigellus Bleuse, 1913 Ille-et-Vilaine
~ehlersi Bernau, 1915; un "N-Spain:Gorbea"
~pyrenaeus Bernau, 1915; un; error for pyrenaicus Lapouge
~simulator Everts, 1915, nec Kraatz, 1876
~viridis Everts, 1918; un Holland:?
~delaunayi Barthe, 1924; un Orne:Domfront
=eques Lapouge, 1925 "Navarra & Dep.Landes"
=pelissieri Darnaud, 1978 Bouches-du-Rhone:Arles:moor area

ssp. graniger Palliardi, 1825 SW,C-Romania:Banat(Mehadia), Bihor; CE-Serbia
~assimilis Duftschmid, 1812, nec Paykull, 1790 "Banat"
~moestus Dejean, 1831, nec Duftschmid, 1812
=nicanor Haury, 1880
=muehlfeldi Géhin, 1885, nec Duftschmid, 1812 ?Retezat Mts.
=szobroniensis Géhin, 1885 ?SE-Romania:Comana
=basalis Beuthin, 1896
=biharicus Reitter, 1896
=rufoscapus Beuthin, 1896
=subgraniger Reitter, 1896 "Klausenburg"
~ater Fleischer, 1898, nec Villers, 1789
=coloripes Fleischer, 1898
=nigripennis Fleischer, 1898
=oxycancellatus Fleischer, 1898
=mazurai Fleischer, 1910 C-Romania:"Retezat:Kimpuluinyag"(=Cimpu lui Neag W of Petroºani)
=disseptus Kolbe, 1913 "Moldova"
~fraternus Kolbe, 1913, nec Csiki, 1906
=insperatus Kolbe, 1913 "Serbia"
=rbanyensis Bernau, 1913 "Banat:Resitzabanya"(=Reºita)
=resiczabanyensis Csiki, 1913; nom.pro rbanyensis Bernau
=romaniensis Kolbe, 1913 "Romania"
=geta Lapouge, 1925; nom.pro szobroniensis Géhin
~pompilii Savulescu, 1992; un Banat:Topenia valley(=ca.24km NE Baile Herculane)
=bibanensis Lie, 1994 Banat:Semenic Mts.:Biban valley(=ca.30km S Caransebeº)

ssp. excisus Dejean, 1826 Austria("Wien:Kahlen- & Leopoldsberges"), NE-Switzerland, S-Germany, Bohemia, S-Moravia
=verrucosus Heer, 1837 ?Switzerland
~marginatus Letzner, 1849; un
~rufofemoratus Letzner, 1849; un
~viridis Letzner, 1849; un
~affinis Gistl, 1857, nec Schrank, 1781 München
~punctulatus Schaum, 1857, nec Schaller, 1783
~cupreoaeneus Dalla Torre, 1877; un ?"Oberösterreich"
~rubrofemoratus Dalla Torre, 1877; un
~viridiaeneus Dalla Torre, 1877; un
~haematomerus Kraatz, 1879, nec Kraatz, 1878
=letzneri Kraatz, 1879
~rufipes Kraatz, 1879, nec DeGeer, 1774
~femoralis Géhin, 1885, nec Motschulsky, 1865
~sudeticus Schulz, 1901; un
=superior Sokolar, 1907 Oberösterreich:Steyr env.
=brevituberculatus Roubal, 1909 ?"Bohemia"
=interior Sokolar, 1910
~opolanus Bernau, 1910; un C-Bohemia:"Opolan bei Podiebrad"(=Opolany nr.Podìbrady)
=pseudoemarginatus Bernau, 1910
=thuringianus Born, 1911 "Thüringen"
=brdensis Sokolar, 1911 C-Bohemia:Brdy Mts.
~bavaricus Kolbe, 1912, nec Beuthin, 1896 Bayern:?
=spaneyi Kolbe, 1912 S-Germany:?
=transitivus Kolbe, 1912 "Thüringen"
=lapougeanus Langenhan, 1913 "Schweiz"
=planianensis Bernau, 1913 "Planian"(=?C-Bohemia:Plaòany W of Kolín)
~pseudobavaricus Bernau, 1913; un Pfarrkirchen
=danubialis Bernau, 1915 S-Germany:?
=maderi Born, 1915 ?Wechsel Mts.
~tersculptus Fleischer, 1917; un "Böhmerwald"(=S-Bohemia:Šumava Mts.)
=transversus Eidam, 1941 "Steiermark"
~baldianus Mandl, 1955; un
~regressivus Poschinger, 1955; un
~gotzi Mandl, 1964; un Marchauen nr.Marchegg(=betw.Wien/Bratislava)
~hanauensis Niedl, 1964; un SC-Germany:Hessen:Hanau
~kubinyii Mandl, 1964; un Niederösterreich:Waldviertel:Gross-Pertenschlag, Sölling, Kasterslehen, Purgstall env.
~wirthumeri Mandl, 1964; un Oberösterreich:Steyr:Reichraming & Kleiberggipfel nr.Schön-Klaus

ssp. intermedius Dejean, 1826 S-Croatia:Dalmatia:Vrgorac; C,S-Bosnia, Herzegovina
=maximus Haury, 1880 Bosnia, NE-Croatia; ?LNF
=apfelbecki Born, 1904 Bosnia:?
=islamitus Born, 1904 Bosnia:Zepce
=livnensis Born, 1906 SW-Bosnia:Livanjsko Polje
=annisus Kolbe, 1913
=duvnensis Bernau, 1914 Bosnia:Duvno Polje
~zoufali Fleischer, 1922; un

ssp. balcanicus Born, 1899 E-Serbia; Bulgaria("Stara Planina nr.Klisura"); ?Eur.Turkey:"Adrianopel"(=Edirne)
=kocae Born, 1910 "Serbia:Slavonia:?"
~drenskyi Breuning, 1928; un S-Bulgaria:Rodopes("Hassa Korja"); alpine form
=simeoni Eidam, 1941

ssp. mimus Kolbe, 1913 Slovenia; SE-Austria, SW-Hungary("Satoristye")
~nigricornis Dejean, 1826, nec Fabricius, 1787 "S-Styria"
~ziegleri Kraatz, 1883, nec Duftschmid, 1812 ?LNF(unlabelled HT); ?"Styria"
~pseudocancellatus Fleischer, 1898; un "Banat"(error)
=ambicornis Sokolar, 1907; nom.pro nigricornis Dejean
~pustuliger Eidam, 1941; un "Tyrolia:Ampezzo Tal:Schluderbach"
~neoziegleri Deuve, 1991; nom.pro ziegleri Kraatz, nec Géhin, 1876 ?LNF(unlabelled HT); ?"Styria"
=neoziegleri Deuve, 2003; nom.pro ziegleri Kraatz, nec Duft., 1812 ?LNF(unlabelled HT); ?"Styria"

ssp. corpulentus Kraatz, 1880 Croatia, Montenegro:Dalmatian coast; ?LNF
=atroviridulus Müller, 1898 "Zara"(=Zadar)
=novaki Müller, 1898 "Vrana Lake"(=Vransko Lake)
=nudilabrus Müller, 1898 "Zara"(=Zadar)
=schatzmayri Born, 1912 "Pago Island"(=Pag)
~beszedesi Depoli, 1915; un "Abbazia"
=ventricosus Bernau, 1915 "Budua"

ssp. alessiensis Apfelbeck, 1901 N-Albania("Velipoja nr.Alessio")
=hypsobius Apfelbeck, 1918 "Nordalbanian Alps:Vunsaj env., karst, 1100-1800m"

ssp. tibiscinus Csiki, 1905 Hungary(?"Tisa Lowland"), NE-Serbia, E-Austria, W,NC-Slovakia, CW-Romania:Timis
~soproniensis Dejean, 1826; nud "Hungary:Oedenburg"(=Sopron)
=adeptus Kolbe, 1913 ?NW-Hungary
=electus Kolbe, 1913 ?W-Slovakia
=inceptus Kolbe, 1913 ?LNF
=nattereri Kolbe, 1913 "Temesvár"(=Timiºoara)
~tatricus Kolbe, 1913, nec Reitter, 1896 NC-Slovakia:Tatry Mts.
=vogeli Breuning, 1933 NW-Hungary:Sopron
=budensis Csiki, 1946 Budapest env.:?Pilis Massif
=leithaicus Mandl, 1955 Burgenland:Leitha Mts.:?



3.vagans Olivier, 1795 SE-France:Alpes-Maritimes, Var(Frejus); NW-Italy:Liguria
~liguricus Lapouge, 1898, nec Born, 1898 Var:Collobriéres
=matheyi Born, 1917 Alpes-Maritimes:Grasse
~borni Barthe, 1921; un
=ligustinus Csiki, 1927; nom.pro liguricus Lapouge "Dep.Var"
=triglyphicus Ochs, 1965 Var:Collobriéres


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CATHOPLIUS Thomson, 1875; type species Carabus asperatus (Dejean)
species: 1



1.asperatus (Dejean, 1826) Morocco:Atlantic coast; West Sahara


ssp. asperatus (Dejean, 1826) Morocco:Atlantic coast:Larache – Casablanca – Safi - Tensift valley; LNF
=cychrocephalus Fairmaire, 1858 "Morocco"

ssp. stenocephalus Lucas, 1866 CW-Morocco:Tensift valley to Cap Rhir; "Mogador"(=Essaouira)
~laevior Escalera, 1914, nec Krynicki, 1830 "Azib el Arjaiat"
=escalerae Csiki, 1927; nom.pro laevior Escalera "Azib el Arjaiat"
~minor Escalera, 1944, nec Kraatz, 1860 Agadir

ssp. ifniensis (Zarco, 1941) SW-Morocco:Sidi-Ifni to Oued Draa; ?Tiznit env.

ssp. susicus Antoine, 1941 CW-Morocco:Cap Rhir to Oued Sous; Aglou

ssp. aliai Escalera, 1944 SW-Morocco & West Sahara:"Jalúa Tantan a Dj.Uarksis"(=nr.Tan Tan) - El Aaiun - ?Boujdour
=mateui Espanol, 1944 Bir Tan-Tan


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HEMICARABUS Géhin, 1885; type species Carabus nitens Linné
species: 4



1.macleayi Dejean, 1826 E,NE-Siberia; Far East, Korea; NE-China:Heilongjiang

ssp. macleayi Dejean, 1826 E,NE-Siberia("Dauria"); Far East, Korea; NE-China:Heilongjiang
=splendidulus Semenov, 1888 E-Siberia:"Bai von Castries"(=De-Kastri S of Nikolaevsk-na-Amure)
=coreensis Breuning, 1933 N-Korea:?Paik-to-san(=Mt.Paektu-san N of Hyesan); ?Hoiryong

ssp. amanoi (Imura, 2004) N-Japan:Rishiri-to Island:Rishiri-zan



2.nitens Linné, 1758 N,W,C-Europe:SW-France to Scandinavia; British Isles, Ukraine, W-Siberia
=aureus DeGeer, 1774
~marginatus Voet, 1778, nec Linné, 1758
=hookeri Nodier, 1821 Scottland
~aureomicans Letzner, 1850; un
~virescens Letzner, 1850; un
=pulchellus Brüggemann, 1873; nom.pro nitens Linné
~cupreus Westhoff, 1881, nec Linné, 1758
~interruptus Westhoff, 1881, nec Fabricius, 1775
=fennicus Géhin, 1885 ?Finland
~niger Semenov, 1886, nec Linné, 1761 Arkhangelsk
=subnitens Reitter, 1896
~funebris Csiki, 1927; un; nom.pro niger Semenov Arkhangelsk
~willemsi (Culot, 1986); un
=aquitaniensis Mollard, 1996 SW-France: near Arcachon
?bistrum Mollard, 1996; un SW-France: near Arcachon



3.serratus Say, 1825 C,SE-USA; S-Canada; North America(by Lindroth:N.Carolina:Ashville)
=lineatopunctatus Dejean, 1826 North America
~ligatus Kirby, 1837, nec Germar, 1824 Canada
=canadensis Melsheimer, 1853; nom.pro ligatus Kirby Canada
=tatumi Motschulsky, 1865 Hudson Bay
=vegasensis Casey, 1914 New Mexico:Las Vegas



4.tuberculosus Dejean, 1829; nom.pro tuberculatus Fischer S,C-Siberia, Far East, Sakhalin, Korea, Japan; "Altay"
~tuberculatus Fischer, 1827, nec Dejean, 1826 "Altay"
=strophium Fischer, 1842 "Dzhungaria"
=granosus Chaudoir, 1844 LNF
=etholeni Mannerheim, 1849 Irkutsk
~marginicollis Kraatz, 1886, nec Duftschmid, 1812
=miyakei Okhura & Goto, 1960 ?Hokkaido


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CARPATOPHILUS Reitter, 1896; type species Carabus linnaei Panzer
species: 1



1.linnaei Panzer, 1812 C-Europe, NE-Italy, Carpathians

ssp. linnaei Panzer, 1812 C-Europe(Germany:Harz Mts.), Carpathians
~angustatus Panzer, 1812, nec Fabricius, 1787 "Austria"
=scopolii Dejean, 1826 "Wolhynien"
~brunneofemoratus Letzner, 1850; un
~cupreoaureus Letzner, 1850; un
~cupreus Letzner, 1850; un
~nigricornis Letzner, 1850; un
~obscurus Letzner, 1850; un
~versicolor Letzner, 1850; un
=hopffgartenianus Beuthin, 1885 "Transsylvania:Kerzeschora"
=bescidensis Reitter, 1896 "Beskiden"(=N-Moravia:Moravskoslezské Beskydy Mts.)
=ludovicus Dietl, 1896 "Rotenturmpass"(=C-Romania:Turnu Rosu Pass)
=quadraticollis Petri, 1912 Romania:"Schässburg"(=Sighiºoara)
=quinquecostatus Petri, 1912 "Schässburg & Kronsdtadt"(=Romania:Sighiºoara & Braºov)

ssp. macairei Dejean, 1826 S-Carpathians:SW-Romania("Mts.of Banat")

ssp. folgariacus Born, 1913 E-Alps:NE-Italy, Austria(?"Südtirol"); SE-Austria:Burgenland
=transdanubialis Kenyery, 1983 Burgenland:Mt.Geschriebenstein
=tauernensis Mandl, 1985 ?Rottenmanner Tauern


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TOMOCARABUS Reitter, 1896; type species Carabus convexus Fabricius
species: 7



1.bessarabicus Fischer, 1823 S-Ukraine, Krym Peninsula; ?Moldavia; SW-Russia; SW,SC-Siberia

ssp. bessarabicus Fischer, 1823 ?Moldavia("Bessarabia:Bender"); S-Ukraine; Krym Peninsula
~tauricus Roeschke, 1897, nec Bonelli, 1811 Krym Peninsula

ssp. concretus Fischer, 1823 SW-Russia("Sarepta"=lower Volga basin); Caucasus; SW,SC-Siberia
=platyscelis Fischer, 1827 "Siberia"
=melambaphus Fischer, 1832 "Kirgizsteppe"
=steppensis Motschulsky, 1846 "Step.Kirgis., fl.Ischim"(LT)



2.convexus Fabricius, 1775 Europe, British Isles, Balcans, Turkey, Russia, Caucasus, Kazakhstan, SC-Siberia

ssp. convexus Fabricius, 1775 C,W,N-Europe(?Germany), British Isles, Russia, Caucasus, Kazakhstan; SC-Siberia:Barnaul
=aemulus Fischer, 1823 Siberia:?Tobolsk, ?Barnaul
~striolatus Fischer, 1823, nec Fabricius, 1801 "Caucasus:N-slope"
=simplicipennis Dejean, 1826 "Silesia"
~gibbus Brullé, 1835, nec Fabricius, 1794
=turcomannicus Motschulsky, 1850 "Turkmenia, Samarkand"(error)
~niger Letzner, 1850; un
~violaceomarginatus Letzner, 1850; un
~viridimarginatus Letzner, 1850; un
=oblongulus Motschulsky, 1865 "Alp.Cauc."(LT)
~coeruleomarginatus Dalla Torre, 1877; un
~viridimarginatus Dalla Torre, 1877; un
=merklii Hopffgarten, 1878 "Siebenburgen"
~borealis Géhin, 1885, nec Paykull, 1790 "Sibérie"
=paganettii Born, 1905 S-Italy:Calabria:Sta.Eufemia d’Aspromonte
~pyrenaicus Born, 1907, nec Lapouge, 1898 Pyrénées-Atlantiques:Valée d’Aspes
=barnaulensis Born, 1922 Siberia:Barnaul
=henseleri Bodemeyer, 1922 Frankfurt a.M.
=appenninus Depoli, 1927 ?"Appennines:Mt.Tresca"
=longobardus Depoli, 1927 "Lombardia"
=pyrenaeicola Csiki, 1927; nom.pro pyrenaicus Born Pyrénées-Atlantiques:Valée d’Aspes
=antipai Panin, 1942 NE-Romania:Piatra Neamt:Varatec
=pseudogracilior Panin, 1942 S-Romania:?
=bucciarellii Mandl, 1966 Alpi Liguri:N of Savona:Mte.San Giorgio
~confusus Tarrier, 1975; un Aude:Le Linas
=kiskunensis Adam & Merkl, 1986 ?SC-Hungary

ssp. dilatatus Dejean, 1826 S-Europe, Balcans Peninsula, C-Europe:S-Austria; NE-Italy:Trieste env.; ?Carniolia
=hornschuchi Hoppe & Hornschuch, 1825; spn Hohe Tauern:Heiligenblut
=perplexus Schaum, 1857 CE-Greece:"Attika & Parnass"(=Attike & Parnassos Massif)
=weisei Reitter, 1883 Adriatic Sea coast:Croatia, Bosnia; ?LNF
=rufulus Géhin, 1885 "High Balkan Mts."(=Stara Planina Mt.R.:C-part)
~proximatus Reitter, 1896; un "Südbosnien"
=cernagorensis Apfelbeck, 1904 Bosnia:Volujak
=chionophilus Apfelbeck, 1904 Bosnia:?
~latus Depoli, 1925, nec Linné, 1758 NW-Italy:?Livorno env.:Abbazia
~proximatoides Depoli, 1925; un NW-Italy:Livorno env.:Abbazia
=miliusi Breuning, 1927 Adriatic Sea coast(numerous localities):S-Dalmatia, Montenegro, N-Albania
=athesinus Depoli, 1927 NE-Italy:Trentino:?
~reductus Müller, 1930, nec Roeschke, 1896 ?"Dalmatia"

ssp. moreanus Reitter, 1896 Peloponnesos(Taygetos Mts.); ?Euvoia Island

ssp. rhinopterus Hampe, 1852 C,S,E,NW-Turkey, Armenia; LNF(?Armenia)
=salax Motschulsky, 1865 Amasya
=sultandaghensis Breuning, 1966 S-Turkey:Afyon:Sultan Dag:Cay env.
=immodicus Cavazzuti, 2006 NW-Turkey:Bursa:Ayva

ssp. acutangulus Chaudoir, 1879 NE-Turkey:Erzurum, Giresun, Ordu, Trabzon
=mackaensis Korell & Cavazzuti, 1991 Trabzon:upper Degirmen Dere valley(=Macka env.)

ssp. gracilior Géhin, 1885 SE-Bulgaria, Romania, NE-Greece, Eur.Turkey; "Transsylvania"
~pygmaeus Géhin, 1885, nec Fabricius, 1792

ssp. erzeliki Schweiger, 1962 NW,NC-Turkey:Bolu(Abant Mts.:grasslands above forest line), Ankara, Sinop
=abantayaginiensis Schweiger, 1962 Bolu:Abant Mts.(Abantayagi:forest zone)



3.decolor Fischer, 1823 W,C-Caucasus("Iberia"); Armenia; NE-Turkey:Kars
=biseriatus Chaudoir, 1846 "C-Caucasus:Glola"
=marthae Reitter, 1896 "Circassia"



4.marginalis Fabricius, 1794 NE,C-Romania; Mold., Ukr.; SW-Hung.:Nagybajom; NE-Poland("Prussia"); C-Siberia:Barnaul
=chrysochlorus Fischer, 1812 "S-Russia & Siberia"
~decorus Seidlitz, 1891, nec Panzer, 1800 "Siebenburgen:Ratosnya"(=Rastolita; 46:59N/25:02E)



5.rumelicus Chaudoir, 1867 C,E-Turkey, Syria, Lebanon, N-Israel, NW-Iran

ssp. rumelicus Chaudoir, 1867 S,C,E-Turkey; "Rumelia"
~cappadociae Breuning, 1933; un Kayseri("Caesarea")
~guelekensis Breuning, 1964; un Adana:betw.Pozanti/Gülek:Ak Dag
=zaraensis Breuning & Ruspoli, 1970 Sivas:betw.Zara/Koyulhisar:Serefeye env.

ssp. kurdistanicus Eidam, 1931 CE-Turkey:Tunceli(Pülümür), Mus, Tatvan; NW-Iran
=kurdistanicola Deuve, 1990 "Armenia turcica:Wan"(=Van Lake env.)
=likbinensis Lassalle, 2006 W-Iran:Azerbaijan:4km E of Likbin

ssp. syriensis Breuning, 1943 NW-Syria(Antilibanon Mt.R.:Hermon), Lebanon
=libanoticus Breuning, 1943 Lebanon:Djebel Sannin
=mendelssohni Schweiger, 1970 N-Israel:Antilibanon Mt.R.:Hermon env.



6.scabripennis Chaudoir, 1850 Turkey, SW-Georgia

ssp. scabripennis Chaudoir, 1850 E-Turkey:Trabzon, Erzurum; "Armenia"

ssp. maljushenkoensis Breuning, 1967 Georgia:Borzhomi

ssp. ponticola Deuve & Simard, 1977 Giresun:Kümbet Pass, 2000m
~reticulatus Hampe, 1852, nec Fabricius, 1787

ssp. microderus Chaudoir, 1867 SC-Turkey:Antalya, Konya, Mugla; "Anatolia"
=ledereri Gautier des Cottes, 1868 "Taurus"
=gautieri Géhin, 1885 ?"Taurus"
=finikensis Lassalle, 1993 Mugla:Fethiye:Finik

ssp. reductus Roeschke, 1896 NC-Turkey:Amasya
=nocturnus Schweiger, 1969 Amasya:Merzifon
=subreductus Breuning, 1977 "Sankiri"(=Çankiri)

ssp. martaiellus Rapuzzi & Kleinfeld, 2000 NW-Turkey:Bolu:Abant Lake, 1500m

ssp. kocaelicus Cavazzuti, 2006 NW-Turkey:Kocaeli:Kargaköy



7.simardianus Deuve, 1990; inc; ?artifact C-Turkey:Amasya; 1m


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EURYCARABUS Géhin, 1885; type species Carabus numidus Castelnau = Carabus famini numidus Castelnau
species: 2



1.famini Dejean, 1826 Sicily, N-Algeria, Tunisia; N,C-Morocco

ssp. famini Dejean, 1826 Sicily

ssp. numidus Castelnau, 1835 NE,NC-Algeria:sea coast("Bone"); NW-Tunisia:Le Kef, Beja env.
=varvasi Solier, 1835 Algiers env.
=hipponensis Fairmaire, 1858 "Bone & Bougie"(=Bejaia)
=oranensis Fairmaire, 1858 Oran(?error)
=berberus Géhin, 1885 "Bougie"(=Bejaia)

ssp. lucasi Gaubil, 1849; nom.pro rugosus Lucas NE-Algeria:inland:Tebessa, Batna; NE-Tunisia:Tunis env.
~rugosus Lucas, 1849, nec Fabricius, 1775 Algeria
=gerardi Lallemant, 1868 Batna

ssp. maillei Solier, 1835 NW-Algeria & NE-Morocco:Tiaret-Mostaganem-Oujda-Melilla; "Barbaria"
=peletieri Castelnau, 1835 "Oran"
=microcephalus Gistl, 1857 "Barbaria"
=lepeletieri Bedel, 1895; ?nom.pro peletieri Castelnau
=pimelioides Lapouge, 1903 W-Algeria:"Frenda"
~verrucosus Lapouge, 1903, nec Heer, 1837 Algeria:"Quarsenis"

ssp. favieri Fairmaire, 1859 N-Morocco:Rif Mts(Tanger; inexact)
=polychaeta Thomson, 1875 "Caucasus"(error)
=otini Antoine, 1939 N-Morocco:SW of Taza:Tazeka Massif
=grasselyi Antoine, 1941 NE-Morocco:Rif Mts:CE-part(Tizi Ifri)
~tazekensis Antoine, in litt. Tazeka Massif:"Msoun"
~tazaotanus (Cobos, 1961); un Rif Mts:E-part:Jb.Tazaot
~ketamensis (Antoine, 1962); un Rif Mts:C-part:17 km Ketama:betw.Tétouan/Alhucemas:cedar forest
~pardoi (Antoine, 1962); un Rif Mts:E-part:Ixmoart
~saisensis (Kocher, 1965); un steppe plateau N of Fes:Sidi-Beurnoussi

ssp. piraticus Fairmaire, 1880 CW-Morocco:Atlantic coast:Larache - Ribat - Dar el Beida
=mogadoricus Escalera, 1914 "El Kureimat SE of Mogador"(=Atlas Mt.R.foothils SE of Essaouira)
=zaianus Antoine, 1941 Oulmés(=Pays Zaer Zaiane area E of Dar el Beida)
~mamorensis (Kocher, 1964); un Foret de Mamora(=betw.Meknes/Kenitra)
ssp. fezzanus Lapouge, 1915 Morocco:Middle Atlas; "Fes"(inexact)
=itoensis Antoine, 1939 N of Azrou:Ito plateau; betw.El Hajeb/Sefrou
~nebulo Antoine, 1939; un Middle Atlas:Ifrane
~vermiculatus (Kocher, 1965); un Middle Atlas:Aguelmane Sidi-Ali, 2000m
ssp. atlantis Alluaud, 1923 Morocco:High Atlas:N-macroslope(NE-part:Tizi Machou)
=mairei Antoine, 1925 High Atlas:"Dj.Aouldjid"(=Toubkal Massif)
~dakkai (Kocher, 1965); un High Atlas:Agoudim:Masker Massif



2.genei Gené, 1839 Corsica, Sardinia(Limbara Mts.)
=lacrymosus Lapouge, 1899 Corsica:Bastia
=kraussei Born, 1908 Sardinia:Oristano


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NESAEOCARABUS Bedel, 1895; type species Carabus interruptus Dejean = Carabus abbreviatus Brullé
species: 4



1.abbreviatus Brullé, 1835; nom. interruptus Dejean Canarias:Teneriffe:N,C-part; "Madeira"(by error)
~interruptus Dejean, 1831, nec Fabricius, 1775 "Madeira"(error)
~viridimicans Enderlein, 1929, nec Kraatz, 1878 LNF (colour form)



2.coarctatus Brullé, 1838 Canarias:Gran Canaria
~eliasenae Uyttenboogaart, 1937; un LNF (colour form)



3.faustus Brullé, 1838 Canarias:Teneriffe:N-part

ssp. faustus Brullé, 1838 Canarias:Teneriffe:NE,NC-part, Anaga Massif

ssp. cabrerai Enderlein, 1929 Canarias:Teneriffe:NW-part:Teno Massif
=tenoinus Enderlein, 1929 Teneriffe:NW-part:Teno Massif



4.gomerae A.Müller, 2004 Canarias:La Gomera Island:La Atalaya env.


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TMESICARABUS Reitter, 1896; type species Carabus cristoforii Spence
species: 1



1.cristoforii Spence, 1823 Pyrenees:N-Spain, S-France(Hautes-Pyrenées:Pic du Bréveu), Andorra
=nicolasi Reitter, 1888 Pic de Néré
~obtritus Lapouge, 1898, nec Chaudoir, 1857
~rufipes Nicolas, 1898, nec DeGeer, 1774 Pic de Néré
=beuthini Schulz, 1899 "Pyrenees"
~clarissimus Meyer, 1996; un


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CAVAZZUTIOCARABUS Imura, 1998; type species Carabus latreilleanus Csiki
species: 1



1.latreilleanus Csiki, 1927; nom. latreillei Dejean; ?non priority W-Alps:NW-Italy, S-Switzerland:Alpi Pennine; "Alps of Piemonte"
~latreillei Dejean, 1826, nec Fischer, 1822 "Alps of Piemonte"
=bremii Stierlin, 1881; ?un; ?priority
=neglectus Kraatz, 1887
=schilskyi Kraatz, 1887


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ORINOCARABUS Kraatz, 1878; type species Carabus sylvestris Panzer
species: 7



1.alpestris Sturm, 1815 E-Alps:NE-Italy:Alpi Carniche, Dolomiti, Adamello Massif; Slovenia; S-Austria

ssp. alpestris Sturm, 1815 E-Alps:NE-Italy:Alpi Carniche; Dolomiti:N-part; Slovenia; SC,C-Austria(Schneeberg) =olivaceus Géhin, 1876
=illyricus Kraatz, 1878 ?"Illyria"
~hickeri Breuning, 1927; un "Böses Weibele"(=Lienz)
ssp. hoppei Germar, 1824; ?iss of hoppii Germar E-Alps:NE-Italy:Alto Adige; S-Austria:Tirol, Salzburg, Karnten; "Austria"
~hoppii Germar, 1824; ?cos of hoppei Germar
=tyrolensis Kraatz, 1876 Tirol:Zillertaller Alpen:Hocheder
=brevicornis Kraatz, 1878
~puncticollis Kraatz, 1878, nec Paykull, 1798
~niger Schilsky, 1888, nec Linné, 1761 Zirbitzkogel
~viridis Schilsky, 1888, nec Dejean, 1826 Zirbitzkogel
=rottenmannicus Sokolar, 1907 Rottenmanner Tauern:Steinamandl
~confluens Born, 1913, nec Fischer, 1827 Rottenmanner Tauern:Hochschwung
~tauernicus Csiki, 1927; un; nom.pro confluens Born Rottenmanner Tauern:Hochschwung
=latschuranus Born, 1921 Kalkalpen:Latschur

ssp. dolomitanus Mandl, 1956 NE-Italy:Dolomiti:Mt.Plose

ssp. adamellicola Ganglbauer, 1904 NE-Italy:Adamello Massif(Mt.Frerone & Val di Leno), Brenta Massif



2.bertolinii Kraatz, 1878 NE-Italy:Dolomiti("Malga Fornasa"=?Colle di Fornace); Belluno env.
=cavallensis Bernau, 1914 Belluno env.:Mt.Cavallo



3.castanopterus A.& J.B.Villa, 1833 NC-Italy/S-Switzerland:Alpi Orobie; Bernina Massif; "Alps of Lombardia"
=putzeysianus Géhin, 1876; nom.pro putzeysi Thomson "Alpes Helvetiae"(LT)
~putzeysi Thomson, 1875, nec Mors, 1863 "Alpes Helvetiae"(LT)
=lombardus Kraatz, 1878 Lombardia:?
=danieli Reitter, 1896 Alpi Orobie:Val Arigna
=mesolcinus Born, 1906 Switzerland:passo della Forcola
=montarerae Bernau, 1914 Alpi Orobie:Mt.Arera



4.concolor Fabricius, 1792 Italy/Switzerland:Trentino; Alpi Pennine, Alpi Lepontine, Berner Alpen

ssp. concolor Fabricius, 1792 Italy/Switz.:Trentino; Alpi Pennine, Berner Alpen; "Germany:Halsatia"(=?Alsace; error)
~alpinus Dejean, 1826, nec Paykull, 1790 "Alps of Italy & Switzerland"(=Alpi Pennine)
=amplicollis A.& J.B.Villa, 1844 "Lombardische Alpen"
=bernhardinus Kraatz, 1878 Italy/Switzerland:Gr.St.Bernhard Pass
=mimethes Kraatz, 1878 Mte.Rosa
~?minor Kraatz, 1878, nec Kraatz, 1860
=bernensis Born, 1895 Switzerland:?"Berge links des Haslitales von der Grimsel bis zum Urbachtal"

ssp. lepontinus Born, 1908 NW-Italy:Alpi Lepontine



5.fairmairei Thomson, 1875 W-Italy:Alpi Cozie, Alpi Graie; SE-France:Savoie

ssp. fairmairei Thomson, 1875 NW-Italy:Alpi Graie:Val Orco-Val Stura-Dora Riparia; "Pyrenees"(error)
=cenisius Kraatz, 1878 Mt.Cenis Massif
=sellae Kraatz, 1878 LNF
=sturensis Born, 1899 Alpi Graie:Colle della Pietra Scritta
~mauriennensis Carret, 1904; un
=nigrocyanescens Carret, 1904

ssp. ceresiacus Born, 1896 NW-Italy:Alpi Graie:Alpe Piane:Ceres env.

ssp. iseranicus Deuve & Simard, 1977 SE-France:Savoie:Iseran Massif(Col de l’Iseran, 2500m)

ssp. baudii Kraatz, 1878 W-Italy:Alpi Cozie:upper Pellice & Po valley(Mte.Viso)
=fairmairei: auct., non Thomson

ssp. stecki Born, 1902 SE-France, NW-Italy:Alpi Cozie:Col Longet, Col de Mary

ssp. pelvicus Cavazzuti, 1973 NW-Italy:Val Varaita(Mt.Pelvo d’Elva)

ssp. heteromorphus Daniel, 1896 NW-Italy:Alpi Graie:Monte Rosa Massif(Col Nuovo)

ssp. fenestrellanus Beuthin, 1899 NE-Italy:Alpi Cozie:Col di Finestre, Albergian, Sestrierres
~cavazzutii Raynaud, 1971; un



6.pedemontanus Ganglbauer, 1892 SE-France & NW-Italy:Alpes-Maritimes, Alpi Liguri, Alpi Cozie

ssp. pedemontanus Ganglbauer, 1892 NW-Italy:Alpi Liguri(Mt.Jurin:Val Pesio; LT); Alpi Marittime
=tendanus Born, 1898 Alpi Marittime:Col di Tenda env.

ssp. omensis Born, 1901 NW-Italy:Alpi Marittime:Val Maira-Val Stura; Mt.Omo & Mt.Baudia

ssp. bisioi Cavazzuti & Meli, 1999 NW-Italy:Alpi Cozie:betw. Maira/Varaita valley(Colle di Bellino)

ssp. maurinensis Deuve, 2002 SE-France:Alp.Hte.Prov.:Alpes Cottiennes:Maurin(=Col de Sautron, ~44:29N/06:53E)

ssp. germanae Casale & Cavazzuti, 1977 NW-Italy:Alpi Marittime:Cuneo:Mt.Saccarello; Mt.Fronte

ssp. raynaudianus Deuve & Simard, 1977 SE-France:Alpes-Maritimes:Colle Lunga

ssp. vesubianus Deuve, 2002; nom.pro maritimus Schaum SE-France & NW-Italy:Alpes-Maritimes:Val Stura-Col de Tenda; LNF (unlabelled LT)
~maritimus Schaum, 1856, nec Motschulsky, 1850 LNF (unlabelled LT; originally "Seealpe"=Alpes Maritimes)
=putzeysianus: auct., non Géhin



7.sylvestris Panzer, 1793 C-Europe:Alps; Slovenia, Carpathians

ssp. sylvestris Panzer, 1793 C-Europe:C,W-Alps:France, Switzerland; C-Germany(Harz Mts.), Bohemia, Moravia
~concolor: Panzer, 1809, non Fabricius, 1792 ?"Harz"
=nivosus Heer, 1837 "High Alps of Switzerland"
~nivalis Heer, 1841; un; error for nivosus Heer
~aeneus Letzner, 1850; un
~aeruginosus Letzner, 1850; un
~brunneus Letzner, 1850; un
~detritus Letzner, 1850; un
~purpurascens Letzner, 1850; un
~virescens Letzner, 1850; un
=sylvinus Gistl, 1857 Tirol
=thueringianus Schilsky, 1880 "Thüringen"
=raxicola Reitter, 1896 Raxalpe
=silesiacus Reitter, 1896 "Oberschlesien"
~extroversus Lapouge, 1903; un
=hortensoides Sokolar, 1910 Stilfserjoch
=micklitzi Sokolar, 1910 "Alps of Nieder- & Oberösterreich"
~blumenthalianus Mandl, 1963; un Harz:Bruchberg-Hochmoor
=slabai Niedl, 1968 S-Bohemia:Babylon nr.Domažlice
=guyicolasi Deuve & Simard, 1976 CW-Switzerland:Jura:Colle d’Aiguillon

ssp. transsylvanicus Dejean, 1826 Carpathians; "Transsylvanian Alps"
=polonicus Motschulsky, 1850 "Polonia"
~glacialis Miller, 1859, nec Gautier des Cottes, 1859 Tatra Mts.(=Slovakia:Tatry Mts.)
=milleri Thomson, 1875; nom.pro glacialis Miller Tatra Mts. (=Slovakia:Tatry Mts.)
=tatricus Reitter, 1896 ?"Hohe Tatra"(=Slovakia:Vysoké Tatry Mts.)
~ligneus Lapouge, 1903; un
=caraimanicus Marcu, 1934 Romania:?Caraiman Mts.
~giumaleuensis Marcu, 1934; un Romania:?Giumalau Mt.R.

ssp. redtenbacheri Géhin, 1876; nom.pro angustatus Redtenbacher E-Alps:SE-Austria(Schneeberg), Slovenia
~angustatus Redtenbacher, 1849, nec Fabricius, 1787 SE-Austria:Schneeberg
=kolbi Breuning, 1927 Slovenia:"Triglav Massif & Steiner Alpen"(=Kamniške Alpe)

ssp. haberfelneri Ganglbauer, 1892 E-Alps:SE-Germany, NC-Austria(Niederösterreich:Ötscher & Dürrenstein)
~cupreoaeneus Dalla Torre, 1877; un
~nigroaeneus Dalla Torre, 1877; un
~viridiaeneus Dalla Torre, 1877; un
~fallax Sokolar, 1910, nec Olivier, 1795 "E-Central Alps"
=petzianus Born, 1911 Dachstein
~vafer Csiki, 1927, nec Duftschmid, 1812; nom.pro fallax Sokolar "E-Central Alps"
=neovafer Deuve, 2002; nom.pro vafer Csiki "E-Central Alps"


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OREOCARABUS Géhin, 1876; type species Carabus errans Gory = Carabus amplipennis Lapouge
species: 5



1.amplipennis Lapouge, 1924; ?non priority NW-Spain; N-Portugal

ssp. amplipennis Lapouge, 1924; ?non priority NW-Spain; ?"N-Spain"
~errans Gory, 1839, nec Fischer; 1823 Sierra de Caniza
~femoralis Beuthin, 1885, nec Motschulsky, 1865
=luetgensi Beuthin, 1886; nom.pro femoralis Beuthin; ?un;?priority
=pedator Reitter, 1887
=oliveirae Csiki, 1927; nom.pro errans Gory Sierra de Caniza

ssp. getschmanni Lapouge, 1924 NW-Spain:Leon, Cantabrian Mts.

ssp. pseudosteuarti Lapouge, 1924 N-Portugal
~steuarti: Heyden, 1880, non Deyrolle, 1852



2.carinthiacus Sturm, 1815 E-Alps:NE-Italy; S-Austria; C-Austria:Rottm.Tauern; Slovenia(Karavanke:Zelenica), Bosnia
=mollii Dejean, 1826
~borni Csiki, 1905, nec Meyer-Darcis, 1902 Bosnia:?
=moosbruggeri Born, 1913 Rottenmanner Tauern:?
=friaulicus Born, 1914 Mt.Cavallo & Musiè nr.Bergogna
=kendii Csiki, 1926; nom.pro borni Csiki Bosnia:?



3.ghiliani La Ferté-Sénectere, 1847 C-Spain

ssp. ghiliani La Ferté-Sénectere, 1847 C-Spain:Madrid, Segovia:Sra.Guadarrama & Sra.Gredos; Spain

ssp. negrei Breuning, 1966 C-Spain:Salamanca:Sierra de Bejar



5.guadarramus La Ferté-Sénectere, 1847 C,NC,E-Spain(Sierra Guadarrama)
=duerensis Raynaud, 1974 Soria:Molinos de Duero
~sagranus Bra?es, 1985; un SE-Spain:Granada:Sierra de Sagra, Sierra de Segura
~hirschfelderi Bra?es, 1987; unCW-Spain:Salamanca:Navasfrias, Pto de Perales, S.Martin de Trevejo
=andiaensis Forel & Leplat, 1998 NE-Spain:Navarra:Sierra de Andia
=guadalupensis Mollard, 2003 W-Spain:Caceres:betw.Canamero/Berzocana:Sierra de Guadalupe



6.hortensis Linné, 1758 C,N,E-Europe; Scandinavia; Eur.Russia; Balcans

ssp. hortensis Linné, 1758 C,N,E-Europe; Scandinavia; Eur.Russia; Bulgaria; Bosnia, Montenegro
=striatus DeGeer, 1781
=gemmatus Paykull, 1790
~cupreoaureus Dalla Torre, 1877; un
~viridiaureus Dalla Torre, 1877; un
~alternans Kraatz, 1878, nec Fabricius, 1792
=duerckianus Ganglbauer, 1890 München
~kelecsenyi Lapouge, 1903; un
~herzegowinensis Apfelbeck, 1904; un "Herzegovina & Montenegro"
=rhodopensis Apfelbeck, 1904
~sobotkaensis Šulc, 1907; un E-Bohemia:Sobotka
=ostariensis Born, 1912 Croatia:Ostaria
=starygradensis Born, 1912 Bosnia:Starygrad
=pillichi Bernau, 1914 Hungary:Komitat Tolna
~magdeleinei Lapouge, 1922, nec magdelainei Bleuse, 1914"Macedonia:Florina"(=N-Greece:Makedonía:Florina)
=dofleini Csiki, 1927; nom.pro magdeleinei Lapouge "Macedonia:Florina"(=N-Greece:Makedonía:Florina)

ssp. preslii Dejean, 1830 S-Greece:Peloponnesos("Morea"); Greek Islands:Kerkyra, Kefallinia, Zakynthos
~jonicus Apfelbeck, 1904; un "Korfu"(=Kerkyra)
=pecoudellus Deuve, 1998 "Peloponnesos:Panankaikon"(=?Panachaikon Massif SW of Patra)

ssp. neumeyeri Schaum, 1856 Adriatic Sea Coast; NW-Albania; S-Italy:Calabria; C-Greece; "Dalmatia & Montenegro"
=walteri Reitter, 1896 Croatia:"Sutomore"
=huemmleri Beuthin, 1899
=calabrus Fiori, 1903 S-Italy:Calabria
~pindicus Apfelbeck, 1904; un C-Greece(?Pindos Mts.); Albania
~albanicola Breuning, 1964; un "Albania:Llogora"
~hilleri Mandl, 1964; un S-Albania:Reza Kanalit:Logara


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CYTILOCARABUS Reitter, 1896; type species Carabus cribratus Quensel
species: 2



1.cribratus Quensel, 1806 Caucasus; Caucasus Minor; NC-Armenia; Turkey

ssp. cribratus Quensel, 1806 Caucasus; Caucasus Minor; NC-Armenia; E-Turkey:Artvin, Kars, Rize,Trabzon; "South Russia"
~foveolatus Adams, 1817, nec Duftschmid, 1812
~scrobiculatus Fischer, 1822, nec Fabricius, 1801
=orientalis Osculati, 1844 "Ararat"
=remotus Reitter, 1888 "Circassia"
~montisimeretiis Mandl, 1964; un Caucasus Minor:Adzharo-Imeretinsky Mt.R.(=Meskhhetsky Mt.R.)
=farineli (Machard, 1975) NE-Turkey:Trabzon:Soganli Pass N of Bayburt
=bilmenekus Lassalle, 1996 NE-Turkey:Rize:Ovit Pass

ssp. schubertianus Breuning & Ruspoli, 1974 CE-Turkey:Bitlis(Tatvan)
=negreianus Deuve, 1994 Bitlis:Reºadiye

ssp. pulumuricus Cavazzuti, 1994 CE-Turkey:Tunçeli:Pülümür env.(pass N of Pülümür), Ovacik; Bingöl:Karasu Dag

ssp. harambureae Breuning & Ruspoli, 1974 SC-Turkey:Içel:Bolkar Dag(?error)

ssp. maironei Cavazzuti, 1994 NC-Turkey:Tokat:Camlibel Pass betw.Tokat/Yildizeli

ssp. echarouxi Lassalle, 1996 NW-Turkey:Bolu:Koroglu Dag

ssp. porrectangulus Géhin, 1885 E-Turkey:Trabzon, Bayburt, Giresun, Ordu; "Pontic Alps"
=tazieffi Breuning & Ruspoli, 1969 Ordu:Gölköy(pass betw.Gölköy/Mesudiye), Mesudiye, Koyulhisar
=arpaalaniensis Machard, 1985 Ordu:betw.Mesudiye/Koyulhisar:Arpaalani

ssp. ispiriacus Deuve, 1993 NE-Turkey:Erzurum:betw.Ispir/Ikizdere Pass(=betw.Ispir/Ovit Pass)

ssp. staneki Štìrba, 1931 NC-Turkey:Kastamonu:Ilgaz Dag Massif
=kreissli Schweiger, 1964 Kastamonu:Ilgaz Dag Massif

ssp. yaraligozi Battoni, 1982 NC-Turkey:Kastamonu:Yaraligöz

ssp. graziolii Cavazzuti, 1987 NW-Turkey:Bolu:Yedigoller

ssp. mahouxi Ledoux, 1972 CE-Turkey:Van:28km N Baskale

ssp. ciloensis Deuve, 1994 SE-Turkey:Hakkari:Cilo Dagh, 2700m; 1f



2.gemellatus Ménétriés, 1832 N-Iran:Elborz Mt.R.; Azerbaydzhan:Lenkoran; "Mts.of Talysh"
=ingratus Lapouge, 1907 "Elbrus"(error for Elborz Mt.R.)


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PACHYSTUS Motschulsky, 1865; type species Carabus hungaricus Fabricius
species: 6



1.cavernosus Frivaldsky, 1837 S-Balcans, C-Italy

ssp. cavernosus Frivaldsky, 1837 Serbia, Bosnia, Macedonia, Albania, Bulgaria, Romania:Alba; "Rumelia"
=sterbai Breuning, 1936
~cavernosulus Mandl, 1961; un Bulgaria:Stara Planina Mt.R.:Mt.Botev, 2370m

ssp. variolatus Costa, 1839 C-Italy:Abruzze Mts.(Gran Sasso Massif)
=rosti Beuthin, 1885
=seriatofoveatus Reitter, 1896



2.glabratus Paykull, 1790 NW,C,E-Europe; Scandinavia, Kola Peninsula; N-Italy, Romania, Carpathians; ?NW-Siberia

ssp. glabratus Paykull, 1790 NW,C,E-Europe, Scandinavia, Kola Penins.; W-Romania; ?NW-Siberia; "Finland & Dalmatia"
=laevigatus Scriba, 1790; spn
~ater Letzner, 1850; un
~coerulescens Letzner, 1850; un
~virescens Letzner, 1850; un
=extensus Kraatz, 1885 SW-Romania:Orsova
=punctatocostatus Haury, 1885 "Galicia"
=lapponicus Born, 1909 NW-Eur.Russia:"Lapland"
~angustatus Petri, 1912, nec Fabricius, 1787 "Siebenburgen"
=liebleri Bierig, 1918 Ukraine:"Galicia:Brody"
~sericeus Csiki, 1945, nec Forster, 1771

ssp. gibbosus Heyden, 1866 C,SE-Romania("Banat"); Serbia, ?Bosnia, Croatia, Slovenia; SE,SC-Austria
~breviusculus Kraatz, 1885, nec Kraatz, 1876 "Siebenburgen:Kapellenberge"
=carinthianus Born, 1915 "Alpen von Kärnten"
=ottofischeri Bodemeyer, 1928 Kärnten:Koralpe

ssp. latior Born, 1895 N-Italy ("Piemonte")
=subglabratus Reitter, 1896 "Bozen"(=Bolzano)



3.graecus Dejean, 1826 Greece, Bulgaria, Albania, Macedonia, S-Romania, Turkey

ssp. graecus Dejean, 1826 S,C-Greece; "Greek Islands"

ssp. trojanus Dejean, 1826 Andros, Tinos, ?Crete; "Mytilene"(=Lesvos; by error)
=hormuzakii Born, 1907 "Bukowina"(error)

ssp. kykladensis Breuning, 1961 Kyklades:Naxos, Thira, Milos, Paros

ssp. morio Mannerheim, 1830, nec Duftschmid, 1812; nbm Turkey(Kop Dag nr.Bayburt); S,CE-Romania; Bulgaria; Albania, Macedonia, NE-Greece
=carceli Gory, 1833 "Smyrna"(=Izmir)
~olivieri Castelnau, 1835, nec Dejean, 1826 "Orient"
~acuminatus Ménétriés, 1836, nec Olivier, 1790 "Turkey"
~aethiops Cristoforis & Jan, 1837, nec Panzer, 1797 "Bosporus"
=cavernicola Kraatz, 1880 "Dobrudzha"
=thessalonicensis Apfelbeck, 1904 NE-Greece:Thessalonike
~winkleri Breuning, 1933, nec Weyenberg, 1869 S-Albania:"Maj’e Tartarit"
=neowinkleri Deuve, 1991; nom.pro winkleri Breuning S-Albania:"Maj’e Tartarit"
=muglanus Cavazzuti, 2006 SW-Turkey:Mugla:Kafaca

ssp. ganymed Heinz, 1975 E-Turkey:Tatvan env.(Pass W of Tatvan)

ssp. oertzeni Ganglbauer, 1888 Karpathos Island

ssp. irmasanus Heinz & Korge, 1967; nom.pro felix Heinz & Korge S-Turkey:Antalya:Toros Dag:Irmasan Pass
~felix Heinz & Korge, 1967, nec Semenov & Znojko, 1931 Antalya:Toros Dag:Irmasan Pass

ssp. mazzii Cavazzuti, 1992 SC-Turkey:Konya:Ovacik Dag Mts.



4.hungaricus Fabricius, 1792 SE-Europe, Ukraine, S-Russia, E-Caucasus, Altays, Sayans, Siberia

ssp. hungaricus Fabricius, 1792 Hungary, SW-Slovakia, E-Austria, SE-Moravia, SW-Romania; ?"Hungary"
=viennensis Kraatz, 1877 NE-Austria:Wien
~frivaldskyanus Breuning, 1933; un SW-Romania:"Temesvar"(=Timisoara)

ssp. mingens Quensel, 1806 SW-Russia:Stavropol; E-Caucasus, Dagestan(Kisljar)
=vomax Dejean, 1826

ssp. gastridulus Fischer, 1823 Krym Peninsula, S-Ukraine; "Gouv.Taurien"(=Krym)
=maeotis Fischer, 1823 "Asov Sea coast"

ssp. scythus Motschulsky, 1847 Ukraine:Odessa,"Ekaterinoslav"(=Dnepropetrovsk; LT); W-Russia:Voronezh, Saratov

ssp. cribellatus Adams, 1812 Siberia:Ural-Yakutsk; Altays, Sayans; "Ostsiberien"
=perforatus Fischer, 1822 "Ural"
~thoracicus Germar, 1824, nec Thunberg, 1784
=variolaris Ménétriés, 1849 "Kisilkum"(=Kyzyl-kum desert)



5.pisidicus Peyron, 1854 SC-Turkey:Antalya, Konya, Isparta

ssp. pisidicus Peyron, 1854 SC-Turkey:Antalya, Konya, Isparta; "Caraman"(=ca.40km NW Antalya)

ssp. samai Cavazzuti & Rapuzzi, 1998 SC-Turkey:Antalya/Konya:Alacabel Pass, 1850m; Konya:Tarasci, 1850m



6.tamsi Ménétriés, 1832 Armenia, Azerbaydzhan, N-Iran:Gilan; ?E-Turkey

ssp. tamsi Ménétriés, 1832 Armenia, Azerbaydzhan("Talysh"), N-Iran:Gilan; ?E-Turkey

ssp. dashtensis Heinz, 1970 N-Iran:Gilan:S of Langerud:Kelishum, 2000m

ssp. eckerleini Heinz, 1970 N-Iran:Gilan:Masuleh, 1800m


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MESOCARABUS Thomson, 1875; type species Carabus catenulatus: Fabricius, non Scopoli = Carabus problematicus Herbst
species: 5



1.dufouri Dejean, 1829 S-Spain

ssp. dufouri Dejean, 1829 S-Spain:Cordoba, Granada; "S-Spain & Tanger"(?error)
=rosalesi Reiche, 1863 Cordoba
~papillatus Lapouge, 1902; un Gibraltar
~scaber Lapouge, 1902; un Granada
=salebrosus Csiki, 1927; nom.pro scaber Lapouge Granada
~granadensis Bra?es, 1987; un Granada:Purullena

ssp. baguenai Breuning, 1926 SE-Spain:Almeria, Granada, Murcia; ?Almeria:Águilas



2.lusitanicus Fabricius, 1801 Portugal, Spain, SW-France:Pyrénées-Atlantiques

ssp. lusitanicus Fabricius, 1801 Portugal; CW-Spain:Salamanca:CW-part
=egesippei La Ferté-Sénectere, 1847 N-Portugal:"Oporto"(=Porto)
=schaumi Gaubil, 1849 "Portugal"
=hellwigi Schaum, 1862
=descensus Schaufuss, 1871
=vieirae Oliveira, 1875 CW-Portugal:Leiria
=mediotuberculatus Schaufuss, 1882
=sabrosensis Reitter, 1896 N-Portugal:Sabrosa
=?vivesi Jeanne, 1973
~almanzaensis Meyer & Mollard, 2000; nud Leon:S of Almanza
=almanzaensis Meyer & Mollard in Deuve, 2003 Leon:S of Almanza

ssp. castilianus Dejean, 1826 CW-Spain:Salamanca:N-part(Los Santos)
=bolivari Breuning, 1926 Salamanca:S-part(Bejar); Caceres:N-part; Avila:S-part
=fuentei Breuning, 1926 Salamanca:SW-part(Sierra de la Pena de Francia)

ssp. breuningi Csiki, 1927; nom.pro kricheldorffi Breuning NW-Spain:Orense(Sierra de Queija), Zamora
~kricheldorffi Breuning, 1926, nec Le Moult, 1913 Orense:Sierra de Queija

ssp. brevis Dejean, 1826, nec Duftschmid, 1812; nbm C-Spain:Avila, Madrid, Guadalajara, Segovia, Burgos, Soria, Logrono, Cuenca, Teruel
~complanatus Dejean, 1826, nec Linné, 1767 Castrogeriz nr.Burgos
=albarracinus Ganglbauer, 1886 Teruel:Sierra de Albarracin
=aragonicus Ganglbauer, 1886 Zaragoza:Sierra de Moncayo:San Martin de Moncayo
=complantus Morawitz, 1890 ?Avila; ?Burgos:Castrojeriz
~logronicus Breuning, 1926; un Logrono:?
=sorianus (Jeanne, 1972) Soria:Puerto de Oncala, 1450m
~guardoensis Meyer & Mollard, 1996; un Palencia:W of Guardo:Prado de Guzpena
=guardoensis Meyer & Mollard in Deuve, 2003 Palencia:W of Guardo:Prado de Guzpena

ssp. helluo Dejean, 1826 CE-Spain:Alicante, Valencia, Teruel, Castellon; ?"Spain"; ?"near Guadarrama, ?"S.de Albarracin"
=alicantinus Ganglbauer, 1886 "Alicante"

ssp. latus Dejean, 1826, nec Linné, 1758; nbm SW,SC-Spain(Caceres:Navalmoral de la Mata); CW-Portugal
=luczotii Castelnau, 1832
=gougeleti Reiche, 1863 "Malaga & Córdoba"
=leptopus Thomson, 1876
=alcaracinus Kraatz, 1886 Albacete:?betw.Alcaraz/Riopar; ?Ciudad Real:Pozuelo de Calatrava
=portalegrensis Reitter, 1896 CW-Portugal:Portalegre
~molossoides Lassalle, 1984; un Caceres:NE-part:Piornal
~nevadensis Bra?es, 1989; un Granada:Guadix

ssp. antiquus Dejean, 1826 S-Portugal; CW-Spain:Caceres/Badajoz:W-part; "Hispania"
=mertolaensis Breuning, 1975 S-Portugal:betw.Beja/Faro:Mertola

ssp. trabuccarius Fairmaire, 1857NE-Spain:Tarragona, Barcelona; Huesca:NE-part; Pyrenées-Or.:Col de Perthus(?error)
=catalonicus Ganglbauer, 1886 Barcelona
~problematicusoides Tarrier, 1974; un; ?hybBarcelona:Collado d’Estenalles; ?hyb lusitanicus/problematicus
=trypalsi Breuning, 1975 Huesca:Mont Perdu Massif:E-slope:Plan de Trypals, 2000m
~troncedoensis Meyer & Mollard, 1990; nud Huesca:Troncedo N of Barbastro
=benabarrensis Meyer & Mollard, 1996 Huesca:Calvera N of Benabarre; LT
=galardonensis Meyer & Mollard, 1996 Huesca:Sierra de Galardon; LT
=troncedoensis Meyer & Mollard, 1996 Huesca:Troncedo env.(=N of Barbastro; LT)
=leyrensis Meyer & Caubet, 1998 Navarra:Sierra de Leyre



3.macrocephalus Dejean, 1826 NW-Spain:Oviedo; Leon:N-part; "Asturia"; S of Puerto de Pajares:Busolongo(NT)

ssp. macrocephalus Dejean, 1826 NW-Spain:Oviedo; Leon:N-part; "Asturia"; S of Puerto de Pajares:Busolongo(NT)
=joaquini Breuning & Toulgoet, 1974 Leon:N-part:Torrebarrio env.
~amayensis Lassalle, 1984; un Palencia/Burgos:Puerta de Amaya

ssp. cantabricus Chevrolat, 1840 NW-Spain:Pontevedra(Sierra de Caniza), La Coruna, Lugo; Orense, Oviedo & Leon:W-part
=liebei Eidam, 1925 La Coruna
=asturicus Born, 1925 Oviedo:W-part:Los Cabos nr.Pravia
=moroderi Breuning, 1926 Mts.of Leon Province"
~magdalenai Meyer & Mollard, 1996; un León:Pto.de la Magdalena
=magdalenai Meyer & Mollard in Deuve, 2003 León:Pto.de la Magdalena

ssp. brabeus Schaufuss, 1876 NC-Spain:Santander to Pamplona (Guipuzcoa:Mt.Aralar); NW-France:Pyrenées-Atlantiques
=ahascoanus Lapouge, 1924 Navarra:Mt.Ahascoa
=barcelecoanus Lapouge, 1924NW-France:Pyrenées-Atlantiques:"Col Barceleco sur la Rhune"(=Col des Trois Fontaines)
=macrocephaloides (Jeanne, 1972) Navarra(Sierra de Andia:Venta de Zumbelz, 1000m)
~rodriguezi Meyer & Mollard, 1998; un Burgos:10km W Poza de la Sal
=rodriguezianus Meyer & Mollard in Deuve, 2003 Burgos:Puerto de Burgos



4.problematicus Herbst, 1786 C,W,N-Europe, Scandinavia, Iceland, Kola Peninsula

ssp. problematicus Herbst, 1786 Austria, N-Italy, Alps; Moravia, Slovakia; "Österreich"
=catenulatus Duftschmid, 1812, nec Scopoli, 1763
~austriacus Sturm, 1815, nec Fabricius, 1801 "Österreich"(inexact; ?Moravia)
=cyanescens Sturm, 1815 ?"Mts.of Austria"
~azurescens Dalla Torre, 1877; un "Oberösterreich"
~marginatus Dalla Torre, 1877; un "Oberösterreich"
~nigrescens Dalla Torre, 1877; un "Oberösterreich"
=angustior Born, 1885 Lugano env.(Mt.Generoso); SC-Switzerland:Ticino
~quadraticollis Lapouge, 1899; un "Basses-Alpes:Riez"(=Alpes-Hte-Provence)
=clairvillei Lapouge, 1916 Savoie:Col du Petit St.Bernard
~iserensis Breuning, 1933; un Isére:Villars de Lans

ssp. harcyniae Sturm, 1815 C,N-Germ.(Harz); N,C-France, Belg., Holl., Denm., Br.Isles; Bohemia, N-Poland, S-Scandin.
=?dissitus Fischer, 1826 "Bessarabia"(error)
=?beauvoisi Dejean, 1826 "North America"(error)
=?californicus Motschulsky, 1865 "California"(error)
=solutus Oberthür, 1884 N-France:Ft.Lorges
=gallicus Géhin, 1885 "Vogesen"
~tibialis Bleuse, 1896, nec Paykull, 1790 N-France:?
=muelverstedti Reitter, 1896 N-Poland:"W.Preussen:Rosenberg"=?Susz )
=wockei Born, 1898 C-Norway:"Dovrefield"
~arvernus Lapouge, 1902; un Auvergne
=bayeri Lapouge, 1910 Switzerland/Germany:Bodensee
=belgicus Lapouge, 1913 Belgium:"Liége, Luxemburg"
=cychriformis Bleuse, 1913 N-France:?
~veneti Auzat, 1915; un Clermont-Ferrand
~anthracinus Everts, 1918, nec Illiger, 1798 Holland:?
~progressus Lapouge, 1924, nec Semenov, 1888 British Isles:?
=scandinavicus Born, 1926 S-Norway & S-Sweden
=cruris Csiki, 1927; nom.pro tibialis Bleuse N-France:?
=procedens Csiki, 1927; nom.pro progressus Lapouge British Isles:?
=obenbergerianus Breuning, 1927W,S-Bohemia("Preitenstein, Scheiben, Rakovitz"=Neètiny, Šejby nr.Nové Hrady, ?Rakovice )
~lhommei Hoffmann, 1928; un N-France:?
~montanus Venet, 1930; un Auvergne
~relictus Hellén, 1934, nec Semenov, 1898 Russia:Hogland Island(=betw.Helsinki/St.Petersburg)
=alluaudi Colas, 1936
~diluvialis Blumenthal & Nüssler, 1967; un Niedersachsen:Lüneburger Heide
=oceanicus (Jeanne, 1971) Gironde:Le Porge-Océan
~campinus Delacre, 1975; un Belgium:Anvers:Zoersel
~pseudogallicus Delacre, 1975; un Belgium:Brabant:Foret de Soignes
~lovervalensis (Mal, 1981); un Belgium:Hainaut:Bois de Loverval S of Charleroi

ssp. inflatus Kraatz, 1878 SE,SC-France:Alpes-Marit.to Pyren.-Or.; NW-Italy:Liguria; "Marseille, Basses-Alp.& Pyr.-Or."
~brisouti Fauvel, 1863; un Pyrenees-Orientales:Valeé d’Eyne
=solidus Lapouge, 1902 Pyrenees-Orientales:Vernet-les-Bains
~dellabeffae Breuning, 1933; un Alpes-Maritimes:Mt.Mounier
~battonii Mandl, 1966; un Liguria:Savona:Osiglia
=falsinflatus Ochs, 1967
~moderatus (Raynaud, 1971); un Alpes-de-Hte-Provence:Vallée de la Gordolasque:St-Grat
=prunieri Pham, 1978 Pyrenees-Orientales:Puigmal Massif

ssp. planiusculus Haury in Géhin, 1885 C,W-Pyrenees(Hautes-Pyrénées:Gavarnie); SC-France:Tarn; NE-Spain:Barcelona, Gerona
=mayeti Géhin, 1885 "Alpes et Pyrénées"
=mulsanti Géhin, 1885 ?Hautes-Pyrénées
=trapeti Bleuse, 1885 Hautes-Pyrénées:Baréges env.:Montferrat Glacier, Vignemale Massif
~niger Nicolas, 1898, nec Linné, 1761 Hautes-Pyrénées:Pic de Néré
=andorranus Lapouge, 1902 Andorra
=cunii Lapouge, 1903 ?Gerona:Rosas
=bepmalei Lapouge, 1904 "Aragonia:Huesca:Col de Basibé"
=jugicola Lapouge, 1908 Haute-Garonne:Luchon env.
=occitanus Lapouge, 1910 Tarn(Montagnes-Noires) to C-Pyrenees
~impar Lapouge, 1910; un Tarn:Montagnes-Noires
~amplicollis Bleuse, 1913, nec A.& J.B.Villa, 1844
~gracilis Bleuse, 1913, nec Küster, 1846
=xaxarsi Born, 1917 Barcelona:Centellas(LT)
~secundariofilicatus Codina, 1921; un Gerona:Setcasas
=oberthueri Venet, 1930 ?Andorra
~nicolasi Venet, 1932, nec Reitter, 1888 Ariége:Foret de Bosc
~navarrensis Breuning, 1933; un Navarra:Mt.Urculo
=nicolasiellus Breuning, 1934; nom.pro nicolasi Venet Ariége:Foret de Bosc

ssp. feroensis Lapouge, 1910 Faer Islands, Hebrides, Shetland & Orkney Isles
~fairensis Breuning, 1966; un Fair Isle(=betw.Shetland/Orkney Isles)

ssp. holdhausi Born, 1911 C-Romania:Carpathians; ?"Siebenburgen:Ceahlau"

ssp. strandi Born, 1926 N-Scandinavia(N-Norway:Lappland:Faestnigstuen); Kola Peninsula

ssp. islandicus Lindroth, 1968 SE-Iceland:Höfn env.:Hornafjördur



5.riffensis Fairmaire, 1875 W-Morocco:Rif Mts.; "Marokko"
=elephas Putzeys, 1872 Rif Mts.
=dominicae Ruspoli, 1976 betw.Chechauen/Ketama(=Rif:Foret de Ketama)


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PLATYCARABUS Morawitz, 1886; type species Carabus depressus Bonelli
species: 5



1.creutzeri Fabricius, 1801 Slovenia, Croatia; N-Italy & SC-Austria:Alps

ssp. creutzeri Fabricius, 1801 Slovenia; Croatia:Rijeka env., Velebit Mts., Kapela Mts.; "Carniolia"
~cupreoaeneus Dalla Torre, 1877; un
~nigroaeneus Dalla Torre, 1877; un
~obscurus Dalla Torre, 1877; un
~viridiaeneus Dalla Torre, 1877; un
=viridimicans Kraatz, 1878 W-Slovenia("Illyria:Kneza, Krn"); E-Italy:Julijske Alpe; SE-Austria:Karavanke:W-part
=schencki Géhin, 1885
=tschapecki Géhin, 1885
=araneipes Lapouge, 1903 Slovenia:"Oberkrain:Stein"(=Kranj env.:?)
=humilis Bernau, 1911 W-Croatia:"Fuzine env.", Velebit, Kapela Mts.
=krnensis Bernau, 1911 W-Slovenia:Krn
=cohumilis Bernau, 1913 Slovenia:"Krain"(=Kranj env.)
=depolianus Bernau, 1913 Croatia:Risnjak
=longellipticus Bernau, 1913 Velebit Mts.
=subalpinus Bernau, 1913 "Illyria:Tolmin, Julijske Alpe etc."
=parumilis Bernau, 1914 Slovenia:"Krain"(=Kranj env.)
=sonticus Bernau, 1914; nom.pro subalpinus Bernau "Illyria:Tolmin, Julijske Alpe etc."
=stolzi Bernau, 1914 "Steiner Alpen:Grintouc & Karawanken:Bodental"(=Kamnishke Alpe & Karavanke)
=trnovensis Bernau, 1914 W-Slovenia:"Ternowaner Wald"(=Trnovski Gozd)
~heros Lapouge, 1924, nec Fabricius, 1801 "Croatia"(error; ?Villach env.)
=carantanus Depoli, 1925 Villach
=pseudoheros Kenyery, 1983 Slovenia:"Bachergebirge:N-slope:1km S Podvelka, 350m"(=ca.betw.Celje/Zagreb)
=posarinii Rapuzzi, 1999 Slovenia:Koper:Kras Mt.R.:Brkini Mts.:Suhorije env.

ssp. kircheri Germar, 1824 N-Italy:Alpi Orobie-A.Carni; Dolomiti; SC-Austria:Gailtaler Alpen; ?LNF
~fossiger Kraatz, 1896, nec Chaudoir, 1877 LNF
=stellifer Semenov, 1898; nom.pro fossiger Kraatz LNF
=grignensis Born, 1902 Bergamasker Alpen:Mt.Grigna
=latschurensis Born, 1912 Latschur
=tesinensis Bernau, 1913 Pieve di Tesino
=hamilcaris Bernau, 1914 Veneto:Belluno env.- Monti Lessini(Mt.Cavallo, Bosco di Cansiglio)
~fruhstorferi Born, 1920, nec Roeschke, 1900 Mt.Generoso
=fruhstorferianus Csiki, 1927; nom.pro fruhstorferi Born Mt.Generoso
~fuliginosus Hölzel, 1950, nec Panzer, 1809 "S of Drau River:Lienzer Dolomiten; N of Drau River:S of Kals:W slopes of Schober Massif"
=rinaldoi Meggiolaro, 1958 Veneto:Conegliano env.

ssp. baldensis Schaum, 1857 NC-Italy:Monte Baldo Massif(C.Telegrafo); betw.Trento/Lago di Garda
~bondonensis Mandl, 1967; un Cornetto Massif SW of Trento:(Mt.Cima Verde)



2.cychroides Baudi, 1860 N-Italy:W-Alps:Susa env.:Mt.Albergian, Finestre Pass
=federicii Born, 1909 Col di Finestra(=Finestre Pass)



3.depressus Bonelli, 1811, nec Paykull, 1790; nbm N-Italy, S,C-Switzerland, S-Austria & SE-France:Alps

ssp. depressus Bonelli, 1811, nec Paykull, 1790; nbm N-Italy, S,C-Switzerland, S-Austria & SE-France(Alpi Graie:Val Soana:Usseglio)
~bonellii Dejean, 1826, nec Adams, 1817 "Alps of Kärnten"
=peirolerii A.& J.B.Villa, 1832
~intermedius Heer, 1837, nec Dejean, 1826 "Graubünden & Wallis"
=glacialis Gautier des Cottes, 1859 "Wallis"
=heerianus Géhin, 1876; nom.pro intermedius Heer "Graubünden & Wallis"
=grajus Reitter, 1896 Alpi Graie:?
=jurinei Born, 1901 Mt.Blanc, Col Ferret, Gr.St.Bernhard
=pseudobonellii Lapouge, 1903
~subparallelus Carret, 1905; un Val Sessera
~transversicollis Carret, 1905; un
=velleri Marceron, 1937

ssp. lucens Schaum, 1856 NW-Italy & SE-France:Alpi Liguri to Alpi Marittime to Alpi Cozie(Mt.Viso:S-slope)



4.fabricii Panzer, 1812 C,SE-Austria, Switzerland & NE-Italy:Alps; Carpathians

ssp. fabricii Panzer, 1812 C,SE-Austria(Niederöst.:Schneeberg); C,W-Switz.; NE-Italy:Alto Adige; W-Carpath.:Slovakia
=heeri Germar, 1824 "Glarner Alpen"
=bugnionii Chaudoir, 1837 Bern:Faulhorn
~polonicus Schaum, 1856, nec Motschulsky, 1865 Carpathians:?
=linderi Tournier, 1860
=poloniensis Géhin, 1876; nom.pro polonicus Schaum Carpathians:?
~nigrinus Schilsky, 1888, nec Motschulsky, 1865 Niederösterreich:Schneeberg
~virens Schilsky, 1888, nec Müller, 1776 Niederösterreich:Schneeberg
=meschniggi Born, 1920; ?hyb "Ostalpen:Ankogel"; ?hyb fabricii/depressus
~fassatii Smetana, 1955; un C-Slovakia:Nízke Tatry Mts., 1750m
=praeglacialis Mandl, 1981 C-Austria:Steiermark:N of Liezen:Mt.Hochmolbing
=feichtauensis Müller & Sach, 1999 NC-Austria:Oberöst.:Molln:Sengsen Mts.:Feichtaualm: betw.Schöneck/Rotgsol, 1400m

ssp. malachiticus Thomson, 1875 Carpathians:N-Romania:Rodna, Caliman Mts.; W-Ukraine:Hoverla, Chorna Gora Massif; LNF

ssp. koralpicus Sokolar, 1910 E-Karnten:Koralpe
=pehri Lapouge, 1913 Koralpe

ssp. ucrainicus Lazorko, 1951 Carpathians:W-Ukraine:Gorgany Mts.



5.irregularis Fabricius, 1792 C-Europe, Croatia, Bosnia, Carpathians

ssp. irregularis Fabricius, 1792 C-Europe:E-France, Switzerland, Belgium, Germany, Austria, Bohemia; LNF(?Germany)
=regularis Fischer, 1823 "Iberia"(error)
=sculptilis Heer, 1837 France:"Niesen & Saléve"
~nigrescens Letzner, 1850; un
~versicolor Letzner, 1850; un
~virescens Letzner, 1850; un
~cephalotes Sokolar, 1909, nec Linné, 1758 ?"NE-Alps"
=jurassicus Born, 1911 ?"C-Jura Mt.R."
=scheffeli Born, 1911 Schwäbischen Alb
~reitteri Appel, 1916, nec Retowski, 1885 Niederösterreich:Lunz
=bitschnaui Born, 1926 Vorarlberg:Freschen
~denesi Niedl, 1973; un S-Bohemia:Hluboká nad Vltavou

ssp. montandoni Buysson, 1882 Carpathians:Slovakia, SE-Poland, Ukraine, Romania(Rarau Mts.:Cruce)
=peronae Hopffgarten, 1885 Romania:"Walachei:Mt.Negoi"
=brunnipes Beuthin, 1885 Romania:"Walachei:Mt.Negoi"
=brostenensis Jacquet, 1886 NE-Romania:Suceava:Broºteni
=narosnyi Csiki, 1946 "Kom.Ung, Poljana"
~minutulus Mandl, 1955; un "Transsylv.Alps:?"

ssp. ramanus Sokolar, 1909 Slovenia, Croatia, Bosnia("Ivan-Sattel"); E-Montenegro:Bjelasica Mts.; SE-Austria:Koralpe
~bucephalus Kraatz, 1879; un Slovenia("Kranj")
=hayeki Kobmann, 1925 ?"S-Steiermark:Possruck"


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CHAETOCARABUS Thomson, 1875; type species Carabus intricatus Linné
species: 2



1.arcadicus Gistl, 1850 C,S-Greece

ssp. arcadicus Gistl, 1850 C,S-Greece:Olympos Massif, Pindos Mt.R.; ?LNF

=adonis Hampe, 1853 ?"Greece"
~parnassicus Mandl, 1955; un Parnassós Massif:?

ssp. merlini Schaum, 1861 S-Greece:Peloponnesos:Taygetos Mt.R.
~borni Kolbe, 1917, nec Meyer-Darcis, 1902 Taygetos
=borniellus Csiki, 1927; nom.pro borni Kolbe Taygetos



2.intricatus Linné, 1761 C-Europe, Italy, Sicily, Carpathians, Balcans

ssp. intricatus Linné, 1761 C-Europe(Germany & ?Sweden), France, Holl., N-Italy, Carpath.; Yug., N-Greece, Eur.Turkey
=cyaneus Fabricius, 1776
=giganteus Heer, 1837; ?in litt. ?Lugano
~gigas Heer, 1841, nec Creutzer, 1799 ?Lugano
~bicolor Letzner, 1850; un
~cyanescens Letzner, 1850; un
~tricolor Letzner, 1850; un
~versicolor Letzner, 1850; un
~violaceus Letzner, 1850; un
~virescens Letzner, 1850; un
=angustulus Haury, 1876 Banat:Bazias
~obscurus Dalla Torre, 1877; un
=montenegrinus Kraatz, 1878 Montenegro
~chlorizans Dalla Torre, 1879; un
=kaadeni Schaufuss, 1880
=bohemicus Haury, 1881 Bohemia
=liburnicus Haury, 1881 Croatia:?
~minor Haury, 1881, nec Kraatz, 1860
~laticollis Roeschke, 1896, nec Sturm, 1826 "Mark Brandenburg"
=germanicus Semenov, 1898; nec ?; nom.pro laticollis Roeschke "Mark Brandenburg"
=neustrius Lapouge, 1900 Bretagne
~aurulentus Lapouge, 1902; un Bretagne
=ulcerosus Lapouge, 1902 Banat
=roeschkeanus Semenov, 1906; nom.pro germanicus Semenov "Mark Brandenburg"
=eurynotus Lapouge, 1913 S-Romania:Comana
=siegwarti Born, 1914 Locarno
=exemtus Kolbe, 1917 "Agram"
=paniscus Kolbe, 1917 Wienerwald
=petax Kolbe, 1917 "Südtirol:Bozen, Klausen, Gardasee"
=bosniensis Born, 1918 Bosnia, Hercegovina & S-Montenegro
=chionobatus Born, 1918 Makedonia/Kosovo:Shar Dag
=starensis Born, 1918 Bulgaria:"Stara Planina, Sofia env., Vitosha, Klissura, Rila Mts."
=carnicus Depoli, 1926 "Alpi Carniche:Mt.Talm"
=macedonicus Jureèek, 1928 S-Macedonia:Pelister Mts.:"Peristeri nr.Monastir"
~weiratheri Breuning, 1934; un NW-Greece:Kerkyra Island; SW-Albania; "Korfu"(=Kerkyra)
~bucovinensis Marcu, 1934; un N-Romania:?
=marchali Raynaud, 1971
=subrhodopensis Blumenthal, 1976 NE-Greece:Thraki:Silo Massif nr.Essimi
~jauffreti Merceron, 1995; un Alpes-Maritimes:N of Nice:nr. Saint*Blaise:Mt.Cima

ssp. lefebvrei Dejean, 1826 Sicily; S-Italy:Calabria:Aspromonte Massif
=aspromontanus Born, 1918 Aspromonte Massif

ssp. bayardi Solier, 1835 S,C-Italy:Roma env.to Calabria(Napoli)
=silaensis Haury, 1881 La Sila Massif
=leonii Born, 1902 "Basilicata"
=molisensis Born, 1918 Latium:Pagano:nr.Castel di Sangro
=lombardii Luigioni, 1927 Firenze(?introduced)

ssp. krueperi Reitter, 1896 E-Greece:Thessalia:Magnesía:"Volo"(=Vólos), Larisa:Ossa Massif


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HETEROCARABUS Morawitz, 1886; type species Carabus marietti Cristoforis & Jan
species: 1



1.marietti Cristoforis & Jan, 1837 N-Turkey, SE-Bulgaria

ssp. marietti Cristoforis & Jan, 1837 NW-Turkey:Istanbul("Bospor, Alem Dag"- NT), Kirklareli, Bolu; SE-Bulgaria:Akhtopol env.
=ativsengueni Schweiger, 1962 Bolu:Abant Mts.
~petrovitzi Mandl, 1964; un Cankiri:Çamlidere
~pseudomarietti Schweiger, 1964; un Bolu:pass betw.Bolu/Kaynasli
=charetianus Auvray, 1992 European Turkey:Kirklareli:Yildiz daglari S of Demirkoy

ssp. sapancaensis Blumenthal & Breuning, 1967 NW-Turkey:Adapazari:Sapanca env.:Goek Dag

ssp. necopinatus Basquin & Darge, 1986 NW-Turkey:Cankiri:Ismetpaºa

ssp. mouthiezi Lassalle, 1991 NW-Turkey:Zonguldak:Ahmetusta Pass

ssp. ~ormanensis Blumenthal & Breuning, 1967; un NW-Turkey:Bolu:Akcakoca:Kaplandere Dagi; Adapazari:Karapurcek:Karadag Massif

ssp. balboi Cavazzuti, 2006 NW-Turkey:Bolu:Bolu Dagi

ssp. bischoffi Chaudoir, 1848 NC-Turkey:Samsun, Amasya; Ordu:Akkus, Gölköy env.; ?Sinop env.; "Ararat"(error)
~kindermanni Hampe, 1852, nec Waltl, 1838
=terahci Charet, 1994 Samsun:Ladik
=delmastroi Cavazzuti, 2003 Amasya:Borabay

ssp. inexspectatus Basquin & Darge, 1986 NC-Turkey:Sivas:N of Koyulhisar, Zara env.

ssp. hroni Deuve, 1997 NE-Turkey:Giresun:Sebinkarahisar env.

ssp. muchei Breuning, 1961 NW-Turkey:Kastamonu:Ilgaz Dag

ssp. stefaniruspolii Breuning, 1966 NC-Turkey:Sinop:Dranaz Dag

ssp. ornatus Heinz, 1976 NW-Turkey:Kastamonu:Yaraligöz, Catalzeytin env.; ?Sinop:Cangal Dag
=inclarus Darge, 1994 Kastamonu:Oyrak Pass

ssp. akensis Haury, 1889 NW-Turkey:Bursa:Ulu Dag Massif(Ak Dag)
=akensiculus Cavazzuti, 2006iarge, 1994 NW-Turkey:Bilecik:Sögüt


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HYGROCARABUS Thomson, 1875; type species Carabus nodulosus Creutzer = Carabus variolosus nodulosus Creutzer
species: 1



1.variolosus Fabricius, 1787 C-Europe, Bulgaria, Yugoslavia, Carpathians

ssp. variolosus Fabricius, 1787 Carpathians; ?Bohemia; "Transsylvania"
=weigeli Panzer, 1801 "Silesia"
=costulifer Fleischer, 1898 "Bihar, Transsylvania"

ssp. nodulosus Creutzer, 1799 C,E-France, Germany, Switzerland, Austria(Linz env.), CW-Hungary, Yugoslavia
=hydrophilus Reitter, 1896 LNF
=goricianus Born, 1911 Italy/Slovenia:near Görz:Panovic


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CTENOCARABUS Thomson, 1875; type species Carabus galicianus Gory
species: 2



1.galicianus Gory, 1839 NW-Spain:Asturia, León, Galicia(Val Pemache); N-Portugal
=beirensis Born, 1924 CW-Portugal:Coimbra



2.melancholicus Fabricius, 1798 N-Morocco, Spain, SW-France

ssp. melancholicus Fabricius, 1798 N-Morocco:Tanger, Taza, Rabat env.; S-Spain
=rotroui Antoine, 1938 N-Morocco:Tazeka Massif:Bou Idir nr.Taza
=mussardi Antoine, 1951 N-Morocco:"marshes of Fouarrat nr.Port-Lyautey"(=Rabat region)

ssp. costatus Germar, 1824 N,C-Spain; N-Portugal; SW-France; "Mts.of Spain"
=pyrenaeensis Géhin, 1885
=castiliensis Born, 1924 C-Spain:Madrid:Sierra Guadarrama:Cercedilla
~leonensis Breuning, 1964; un NW-Spain:Leon:Busdongo
~cuencaensis Raynaud, 1972; un CE-Spain:Cuenca:Olivarés
~exilis Raynaud, 1972; un C-Spain:Guadalajara:Villanueva de Alcojon

ssp. submeridionalis Breuning, 1975 SW,SC-Spain(Malaga:Estepona); S-Portugal
~tietarensis Bra?es, 1987; un C-Spain:Tietar River
=dehesicola Garcia-Paris & Paris, 1996 Cáceres:Torrejón El Rubio


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SPHODRISTOCARABUS Géhin, 1885; type species Carabus adamsi Adams
species: 16



1.adamsi Adams, 1817 C,E-Caucasus; S-Azerbaydzhan; N-Iran

ssp. adamsi Adams, 1817 C,E-Caucasus:Terek River, Vladikavkaz env.- Telavi env.
=chiragricus Fischer, 1828 "thermal springs of Caucasus"(=Pyatigorsk, Kislovodsk env.)
=eichwaldi Fischer, 1828 "Kaukasus"
=chrysitis Motschulsky, 1839 "Alp.Cauc.:Tuschetia"(LT)
=thomsonianus Géhin, 1876
=subcyaneus Kraatz, 1878, nec Illiger, 1802; nbm C-Caucasus:Kazbegi env.; "?Caucasus"
=porphyrobaphes Ganglbauer, 1887 ?LNF(?E of Vladikavkaz)
=adamsianus Mandl, 1975 "Caucasus"
=svaneticola Deuve, 2002 "Svanetia"
~kabardaensis Deuve, 2002, nec kabardensis Kurnakov, 1970 "Kabarda"
=pseudojanthinus Deuve, 2002 LNF(unlabelled HT)

ssp. hollbergi Mannerheim, 1827 E-Cauc.:Shakh D., Kurush; Azerbaydzhan:Kirovabad, Khanlar, Karabakh Mts.; "Dagestan"(LT)
=separandus Kraatz, 1881 "Elisabethpol"(=Kirovabad; =Gyandzha)
~eichwaldianus Géhin, 1885; ?in litt.
~grusianus Roeschke; in litt.
~separandulus Mandl, 1975; un LNF(unlabelled LT)
=adjikentensis Deuve, 2002 Azerbaydzhan:"Adjikent"(=Adjikend; ca.15km SE Gyandzha); 1m
=khanlarensis Deuve, 2002 Azerbaydzhan:"Helenendorf"(=Khanlar; 9km S Gyandzha)
=tchaikentensis Deuve, 2002 Azerbaydzhan:"Tchaikent"(=Chaykend, ca.17km SE Gyandzha); 1m

ssp. serriae Deuve, 2002 N-Iran:Azerbaydzhan:E-part:Kaleybar env.

ssp. separatus Lapouge, 1907 SE-Azerbaydzhan:Lenkoran; N-Iran:Gilan:Asalom, Masuleh; "Talysh"

ssp. persa Lapouge, 1924 N-Iran:Elborz Mt.R.:Ramsar env. to Sari env.; "W-Elborz"(HT lost); Kelardasht (NT)
~longicollis Mandl, 1955, nec Duftschmid, 1812 "Persia, Elburs"
~sahelae Lassalle, 2002; un N-Iran:Mazandaran:betw.Behshahr/Gorgan:Mts. S of Galugah, 1800m

ssp. korgei Heinz, 1970 N-Iran:Mazenderan:Khalus valley:pass N of Gakhsar(~36:10N/51:19E)
=delirensis Deuve, 2000 N-Iran:Mazendaran:SW Chalus:Delir, 36:21N/51:08E

ssp. haerii Heinz, 1973 N-Iran:Gilan:E of Rudbar:Dare-dasht
=larkhaniensis Deuve, 2001 N-Iran:Gilan:Espeyli env.:Larkhani Mts., 1400 m; betw.Siyahkal/Espeyli; Balarud env.

ssp. bararii Deuve, 2000 N-Iran:Mazendaran:Ramsar env.:Ney Dasht, 300-350m

ssp. samaorum Rapuzzi, 2005 NE-Iran:Golestan:Gorgan env.(20km S Gorgan:Nahar Horan)



2.biroi Csiki, 1927; nom.pro kindermanni Chaudoir NC-Turkey

ssp. biroi Ménétriés, 1832 NC-Turkey:Samsun:Kavak env.; Amasya
~kindermanni Chaudoir, 1850, nec Waltl, 1838 Amasya

ssp. iorbi Deuve & Charet, 1998 NC-Turkey:Samsun:W of Ladik

ssp. leonardoi Deuve, 2005 NC-Turkey:Sinop:30km E Ayancik; 1f



3.bohemani Ménétriés, 1832 Azerbaydzhan/Iran borders region; N-Iran:Gilan, Mazandaran

ssp. bohemani Ménétriés, 1832 Azerbaydzhan/Iran borders region:Lenkoran, Ardabil env.; "Mts.of Talysh"
=yvdouarini Deuve & Doline, 2003 Azerbaydzhan:Lenkoran anv.:(Hirkansky Les, Astara, Masilli)

ssp. esperanzae Heinz, 1970 N-Iran:Gilan:NW of Safid valley:Masuleh, Asalam

ssp. esfandiarii Heinz, 1973 N-Iran:Mazandaran:Sardab-rud valley:Rudbarak, S of Chalus:Kelardasht

ssp. morvani Morvan, 1976 N-Iran:Mazendaran:Elborz Mt.R.:Chorteh, 1800m(=SW of Ramsar)

ssp. rosicolor Deuve, 2001 N-Iran:Gilan:Espeyli env.:Larkhani Mts., 1400 m



4.coruhnehriensis Cavazzuti, 1990 NE-Turkey:Artvin:Coruh valley:25-30km W Yusufeli; ?Rize:Ovit Pass:N-slope



5.elegantulus Motschulsky, 1850 N-Iran:E-Elborz Mt.R.; Iran/Turkmenistan:Kopet Dag Mt.R.

ssp. elegantulus Motschulsky, 1850 N-Iran:E-Elborz:Gorgan env.(="Astrabad"), Bojnurd; Iran/Turkmenistan:Kopet Dag Mt.R.
=astrabadensis Semenov, 1903 "Mts. of Astrabad Province"(=Gorgan)
=parthicus Lapouge, 1924 "Budschnurd Mts."(=Bojnurd env.)

ssp. sarafrazii Deuve, 2000 N-Iran:Mazandaran:Sari:Kiyasar:Vesmine Mts.(Kiyasar:36:14N/53:22E)

ssp. dinae Rapuzzi, 2005 NE-Iran:Golestan:S of Gaz, 36:39N/53:58E



6.enigmaticus Heinz, 1980 NE-Turkey:Erzurum, Artvin, Kars

ssp. enigmaticus Heinz, 1980 NE-Turkey:Erzurum:Ispir env.(10km NW Ispir, 1500-1700m)

ssp. reiseri Heinz, 1980 NE-Turkey:Artvin:NW of Yusufeli:Yaylalar, Altiparmak env.

ssp. exiliatus Cavazzuti, 1990 NE-Turkey:Artvin:Kiliçkaya env.:Deve Dag Mts.:1800-2000m

ssp. citamgineus Lassalle & Charet, 1992 NE-Turkey:Kars:Sarikamis, Göle
=bechteli Charet, 1996 Kars:Göle



7.georgia Cavazzuti, 1984 CE-Turkey:Tunceli env.(10km NE of Tunceli)
=hardei Heinz, 1990 CE-Turkey:Tunceli:Kötek-Yayla, Gozen env.



8.giachinoi Cavazzuti, 1991 NE-Turkey:Gümüshane:NE of Siran:Tersundagi Pass, 1800m; 4f



9.gilnickii Deyrolle, 1869 NE-Turkey:Trabzon, Rize, Giresun, Erzurum

ssp. gilnickii Deyrolle, 1869 NE-Turkey:Trabzon env.:Macka, Zigana Pass, Soganli Pass, Ovit Pass
=giulielli Thomson, 1875
=acutus Lapouge, 1904 Trabzon:"Kolat Dag"(=Mts.nr.Zigana Pass)
~viridifossulatipennis Breuning, 1964; un Trabzon:Balabandag W of Macka
~brignolii Breuning & Ruspoli, 1970; un Trabzon:"Kolat Dag"(=Mts.nr.Zigana Pass)
=yakalamakus Lassalle, 1993 NE-Turkey:Giresun:S of Dogankent River:steppes W of Kürtün
=ovoideus Machard, 1994 Trabzon:Soganli Pass S of Caykara
=pseudoelongatus Machard, 1994 Rize:betw.Sivrikaya/Ovit Pass
=lavezorum Darge, 1994 Rize:Ovit Pass:N-slope

ssp. hasandaghensis Darge & Basquin, 1990 NE-Turkey:Erzurum:N of Ispir:Hasan Dag

ssp. brunneicupreus Deuve, 1994 NE-Turkey:Rize:Çat



10.heinzi Breuning, 1964 NE-Turkey:Giresun, Ordu

ssp. heinzi Breuning, 1964 NE-Turkey:Giresun:Egribel Pass, Kulakkaya, Tamdere env.

ssp. mateuellus Deuve, 2003 NE-Turkey:Ordu:Gölköy env.(pass betw.Mesudiye/Gölköy)
~matteui Breuning & Ruspoli, 1970; un NE-Turkey:Ordu:Gölköy env.(pass betw.Mesudiye/Gölköy)



11.kurdicus Heinz, 1975 CE,SE-Turkey:Bitlis, Bingöl, Siirt

ssp. kurdicus Heinz, 1975 CE-Turkey:Bitlis:Tatvan, Resadiye env.
~schuberti Breuning, 1970, nec Breuning, 1968 Bitlis:Tatvan env.
=basquini Darge, 1990 Bitlis:Resadiye/Gevas:Alacabük Dag, 1770m

ssp. machardianus Machard, 1986 CE-Turkey:Bingöl env.; 2f

ssp. kurdaorum Lassalle, 1992 SE-Turkey:Siirt:Yolbilen



12.macrogonus Chaudoir, 1847 N-Turkey:Adapazari to Artvin
ssp. macrogonus Chaudoir, 1847 NE-Turkey:Trabzon env., Macka valley, Zigana Pass; Giresun; LNF
=theophilei Deyrolle, 1869 Trabzon env.
=kolatensis Lapouge, 1924 "Kolat Dagh nr.Trabzon"(=Mts.nr.Zigana Pass)
=koselerensis Machard, 1991 Giresun:betw.Tirebolu/Köseler
=giresunensis Deuve, 1992 "Anatolia, Giresun"(exactly:?Balaban Dag=Mts.SW of Yavuzkemal)
=lemeslei Machard, 1995 Trabzon:S of Caykara:Köknar, 1800m

ssp. pseudokolatensis Heinz & Korge, 1967 NE-Turkey:Artvin:Kackar Dag:N-slope; Rize:Ilica, Ardesen valley, Ikizdere
=rizensis Ledoux, 1975 Rize:S of Pazar
=michaelae Darge, 1994 Rize:Ovit Pass:N-slope

ssp. azaleae Cavazzuti, 1986 NE-Turkey:Ordu:40km SE Ordu:Cambaºi(=E of Topcam)

ssp. resplendens Breuning, 1934 NC-Turkey:Amasya:Borabay; Samsun:Ayvacik, Carsamba, Ladik; Ordu:Ünye
=chambruni Breuning & Ruspoli, 1970 Samsun:Akpinar nr.Carsamba
=cavazzutianus Casale, 1973 Ordu:S of Ünye
=zanderi Battoni & Blumenthal, 1973 Samsun:Ladik
~eiselti Mandl, 1975; un Ordu:betw.Akkus/Ünye

ssp. akkusensis Breuning & Ruspoli, 1970 NE-Turkey:Ordu:Akkus, Gulebi, Ulubey env.; Tokat:Niksar
~akkusensis Breuning, 1964; un Ordu:Akkus
~ingloriosus Breuning & Ruspoli, 1970; un Ordu:Akkus
=guergentepeensis Blumenthal, 1975 Ordu:betw.Gölköy/Ulubey:Gurgentepe Pass
=lilianae Casale & Cavazzuti, 1976 Ordu:Gölköy
~daumani Breuning & Ruspoli, 1976; un Ordu:Ulubey

ssp. simonianus Breuning & Ruspoli, 1970 NE-Turkey:Ordu:betw.Mesudiye/Koyulhisar; Gölköy, Mahmudiye Sivas:Koyulhisar
~pseudosimonianus Mandl, 1975; un Ordu:Gölköy

ssp. diogenei Cavazzuti, 1991 NC-Turkey:Sinop:Gokcebel, Incemeydan, Hatip, Buyukduz, Turkeli

ssp. schweigerinae Schweiger, 1962 NW-Turkey:Bolu(Abant Lake)
=goettingi Schweiger, 1964 Bolu:Abantayagi
=pseudoschweigerinae Schweiger, 1964 Bolu:Abantayagi

ssp. drannadaghensis Breuning, 1964 NW-Turkey:Kastamonu:Ilgaz, Yaraligöz; Sinop:Dranaz Dag, Cangal Dag
=clientulus Schweiger, 1969 Istanbul City (?introduced)
~mollioni Breuning & Ruspoli, 1976; un Sinop:Cangal Dag 40 km W Sinop:S of Ayancik

ssp. keltepensis Vigna-Taglianti & Bruschi, 1976 NW-Turkey:Adapazari:Sapanca env.; Kocaeli:Goek Dag nr.Izmit



13.pavesii Cavazzuti, 1992 W-Turkey:Izmir:Bornova; 1m; probably introduced macrogonus-specimen



14.scovitzi Faldermann, 1835; cos SW-Georgia:Batumi, Makharadze env; NE-Turkey:Artvin, Rize

ssp. scovitzi Faldermann, 1835; cos SW-Georgia:Batumi, Makharadze env.; LNF(?Batumi env.)
~sovitzi Faldermann, 1835; ?iss

ssp. otingoensis Deuve, 1993 NE-Turkey:"Artvin:Karckal-Otingo"(=Karcal Dagi:Balci env.)

ssp. hedini Heinz, 1973 NE-Turkey:Rize & Artvin:Ayder, Yaylalar env.(Ardesen valley:Kavran-yayla)

ssp. bouilloni Breuning, 1976 NE-Turkey:Artvin:Murgul nr.Borçka

ssp. czipkai Heinz, 1980 NE-Turkey:Artvin:Artvin env.(forests above Artvin, 1600-1800m)

ssp. sarigoelensis Deuve & Charet, 1992 NE-Turkey:Artvin:35km W Sarigol



15.tokatensis Roeschke, 1898 NC,CE-Turkey:Tokat, Gümushane, Tunceli, Erzurum

ssp. tokatensis Roeschke, 1898 NC-Turkey:W of Tokat:Topcam Dag Massif; Tokat

ssp. almushensis Mandl, 1975 NC-Turkey:E of Tokat:Almus env.; Yildiz Daglari; Sivas:Koyulhisar

ssp. kosensis Cavazzuti, 1984CE-Turkey:Gümüshane:Köse(pass N of Köse), Siran, Kelkit, Torul env.

ssp. kirikensis Staven, 1995 CE-Turkey:Erzurum:W of Kirik, SE of Kirik

ssp. simonae Cavazzuti, 1984 CE-Turkey:Tunceli:Munzur Daglari:Pülümür, Ovacik

ssp. karasudominus Cavazzuti, 1991 CE-Turkey:Erzurum:Karasu Daglari:Cat env.; 2f



16.varians Fischer, 1827 Armenia, Georgia; SW,SC-Caucasus; NE-Turkey

ssp. varians Fischer, 1827 Armenia:Sevan, Dilidzhan, Yerevan; Georgia:Tbilisi, Somkhet Mt.R.; "Caucaso cisalpino"
=sieversi Ganglbauer, 1887 Caucasus Minor:Trialetsky Mt.R., Bakuriani; "Manglis nr.Tiflis"(=Tbilisi env.)
~praevarians Mandl, 1975; un "Somchetia"
~somcheticus Mandl, 1975; un Armenia:Dilidzhan

ssp. armeniacus Mannerheim, 1830 NE-Turkey:Erzurum(Kop Dag S of Trabzon); Artvin; Georgia:Caucasus Minor, C-Caucasus
=incatenatus Mannerheim, 1830 Kop Dag S of Trabzon
~confusus Fischer, in litt.
=fulminans Kraatz, 1878 ?Borzhomi
=subincatenatus Kraatz, 1878 ?Borzhomi
=repletus Reitter, 1884 SC-Caucasus:upper Inguri & Rioni River; "Svanetia"
~mutabilis Géhin, 1885; un
=korbianus Ganglbauer, 1887 S-Georgia:Batumi env.; NE-Turkey:Artvin:Hopa
=laevilineatus Ganglbauer, 1887 SC-Caucasus:Kodorsky Mt.R., Egrissky Mt.R., Tkvarcheli, Zugdidi env.
=decoloratus Reitter, 1896 ?"Svanetia"
~pseudoarmeniacus Breuning, 1935; un S-Georgia:"Achalzich"(=Akhaltsikhe)
~voriseki Novotný, 1975; un ?Trialetsky Mt.R.:Mt.Ckhra-Ckkhara
=arsiyandagensis Machard, 1985 Kars:NW of Hanak:Arsiyan Dag
=sarikamisensis Cavazzuti, 1986 Kars:Sarikamis
=robichei Auvray, 1994 Artvin:Zeytinlik

ssp. scintillus Reitter, 1884 C-Caucasus:S-slope:"Svanetia"(alpine form)

ssp. janthinus Ganglbauer, 1887, nec Duftschmid, 1812; nbm W-Caucasus:Gagrinsky Mt.R.-Tuapse; Maykop env.; Slavyansk:Krasny Les; Mt.Achishkho(LT)
~ciscaucasicus Mandl, 1975; un Dagomys
~dvorshaki Mandl, 1975; un SW-Caucasus:Pitsunda env., Adler; Bzybsky Mt.R.
~manfredschmidi Mandl, 1975; un Maykop env:Suntuk

ssp. rugatus Breuning, 1934 W-Caucasus:Sukhumi, Novy Afon, Tsebelda, Amtkel, Atsgara, Abkhazsky Mt.R.(Marukh Pass)
~novotnyorum Mandl, 1975; un Sukhumi

ssp. alagoensoides Mandl, 1975 W-Armenia:Mt.Aragats; "Araxestal"(=Araks valley)
~alagoensis Reitter, in litt.
~alagoensoides Mandl, 1955; un "Araxestal"(=Araks valley)

ssp. blumenthaliensis Heinz & Korge, 1967 NE-Turkey:Rize:Kackar Dagi:N-slope(Ilica, Ardesen valley)

ssp. basilewskianus Breuning & Ruspoli, 1970 Artvin:Hopa env.(Ciftekoprüler betw.Hopa/Borcka); ?Ogdem env.
=aabadensis Lassalle, 1991 NE-Turkey:Artvin:Kafkasor nr.Artvin, 1800m

ssp. aequicostalis Staven & Heinz, 1995 NE-Turkey:NE of Artvin:Dalis Dag

ssp. rotundithorax Deuve, 2003; nom. pro rotundicollis Mandl NE-Turkey:Artvin:Karçal Dagi Massif; ?Tunceli; Kartsch Chal(=Karçal Dagi)
~rotundicollis Mandl, 1955, nec Marsham, 1802 Kartsch Chal(=Karçal Dagi)


BACK



OXYCARABUS Semenov, 1898; type species Carabus saphyrinus Cristoforis & Jan
species: 1



1.saphyrinus Cristoforis & Jan, 1837 N-Turkey:Istanbul to Rize

ssp. saphyrinus Cristoforis & Jan, 1837 NE,NC-Turkey:Trabzon, Giresun, Rize, Samsun, Ordu; "Trabzon"(NT)
=wagneri Küster, 1846
~artusi Schweiger, 1964; un Trabzon:coast region
~ruspolii Breuning, 1964; un Ordu:Akkus, Gölköy, Ulubey
=besanconi Breuning & Ruspoli, 1970 Samsun:Carsamba(Akpinar nr.Carsamba), Ayvacik, Kavak; Amasya:Akdag env.
=juengeri Breuning & Ruspoli, 1970; emend.pro yungeri Breuning & Rusp. Ordu:Mesudiye env.(betw.Mesudiye/Koyulhisar)
~yungeri Breuning & Ruspoli, 1970; un; ios
~pseudowagneri Mandl, 1973; un Trabzon:?Zigana Pass

ssp. pseudosaphyrinus Schweiger, 1964 NW,NC-Turkey:Bolu(Abant Mts.:deciduous forest), Kastamonu, Sinop
~pseudosaphyrinus Schweiger, 1962; un Bolu:Abant Mts.:deciduous forest
~boluensis Schweiger, 1964; un Bolu:Abant Mts.:coniferous forest
~korgei Heinz & Korge, 1964; un
=kainashensis Schweiger, 1964 Bolu:pass betw.Bolu/Kaynasli
~rabaronianus Breuning, 1964; unSinop:Dranaz Dag; Kastamonu:Inebolu, Yaraligöz
~ilgazdagicus Blumenthal & Breuning, 1967; un Kastamonu:Ilgaz Dag
~mengenensis Blumenthal & Breuning, 1967; un Bolu:N of Mengen
~yenicensis Lassalle, 1998; unZonguldak:Yenice env., 800m

ssp. pirithous Reitter, 1896 NW-Turkey:Kocaeli:Izmit env.; Adapazari; Alem Dag E of Istanbul
~sapancae Blumenthal & Breuning, 1967; un Adapazari:Sapanca, Goek Dag

ssp. pseudopirithous Mandl, 1965 NW-Turkey:Adapazari:Karapurcek:Karadag Massif; Bolu:Akcakoca; "Caucasus"(error)

ssp. notabilis Roeschke, 1898 NW-Turkey:Bursa:Bursa env., Ulu Dag Massif
=kenyeryi Schweiger, 1964 Bursa:Ulu Dag Massif
~akdagicola Mandl, 1973; un Bursa:Ak Dag(=Ulu Dag) Massif; alpine form


BACK



PROCRUSTES Bonelli, 1809; type species Carabus coriaceus Linné
species: 10



1.anatolicus Chaudoir, 1857 Cyprus, SW-Turkey:Antalya, Mersin

ssp. anatolicus Chaudoir, 1857 Cyprus
=cypricus Roeschke, 1915 Cyprus

ssp. lycicus Breuning, 1934 SW-Turkey:Antalya env.("Lycian Taurus:Adalia"); Mersin, Isparta
=resslianus Mandl, 1963 SW-Turkey:Mersin:Silifke
=gueneycensis Èatloš & Moravec, 1999 SW-Turkey:Isparta:Davraz Dag Mts.:Güneyce env.



2.banoni Dejean, 1829 Crete Island; Dia Island; Gaudos Island
=taborskyi (Maøan, 1948)



3.chevrolati Cristoforis & Jan, 1837 NW,C,E-Turkey; SW-Georgia; ?Armenia

ssp. chevrolati Cristoforis & Jan, 1837 C-Turkey:Amasya(NT), Ankara, Kayseri, Yozgat, Adana, Tokat
~assimilis Cristoforis & Jan, 1837, nec Paykull, 1790 Amasya(NT)
~persimilis Csiki, 1927; un; nom.pro assimilis Cristoforis & Jan Tokat(NT)

ssp. thirki Chaudoir, 1857 Black Sea coast:Kastamonu to Artvin; "Ararat"(error)
~wiedmanni Cristoforis & Jan, 1837, nec wiedemanni Ménétriés, 1836 Mts.S of Sinop(NT)
=surejae Csiki, 1927; nom.pro wiedmanni Cristoforis & Jan Mts.S of Sinop(NT)
~ilgazdaghensis Heinz & Korge, 1964; un Ilgazdag Massif
=dadayanus Schweiger, 1964 "Baumschule Gölköy etwa 20km N von Kastamonu"
~golkoyensis Schweiger, 1964; un "Baumschule Gölköy etwa 10km N von Kastamonu"
~ilgazdagensis Schweiger, 1964; un Ilgazdag Massif:fir forest zone
=chauveloti Breuning & Ruspoli, 1977 Artvin:Hopa
=esmeraldae Breuning & Ruspoli, 1977 S of Samsun
=portei Breuning & Ruspoli, 1977 Ordu:Gölköy
=rodriguezi Breuning & Ruspoli, 1977 Samsun:Akpinar nr.Carsamba:left Yesilirmak River bank

ssp. korbi Breuning, 1928 NE-Turkey:Kars, Artvin; S-Georgia:Akhazlikh; ?Armenia; ?"Transcaucaucasia"

ssp. internatus Heinz & Korge, 1964 NW-Turkey:Bolu(Abant Mts.), Zonguldak; ?Cankiri
~cifcandaghensis Blumenthal & Breuning, 1967; un Ankara/Bolu:Cifcan Dag
=salignicola Schweiger, 1969 Bolu env.:steepe zone

ssp. vanensis Machard, 1985 CE-Turkey:Bitlis:Resadiye, Tatvan; Bingol env.



4.clypeatus Adams, 1817 Caucasus, W-Georgia, Armenia; NE-Turkey:Artvin, N-Iran

ssp. clypeatus Adams, 1817 C,E-Caucasus:Terek River, Dagestan, N-Iran:Kaleybar (~38:52/47:02)
=rectoimpressus Motschulsky, 1850 "Armenia"(LT; ?inexact)
~caucasicus Géhin, 1883, nec Adams, 1817 "Caucasus"

ssp. kurnakovi Kryzhanovskij, 1968 W-Caucasus, W-Georgia, Armenia(Bakuriani), NE-Turkey:Artvin:Karckal Dagi
~fischeri Faldermann, 1835, nec Fischer, 1823
~abchasicus: auct., non Motschulsky



5.coriaceus Linné, 1758 Europe:France to European Russia; Balcans, Carpathians, Turkey, NW-Syria

ssp. coriaceus Linné, 1758 C,W,NE-Europe:France to European Russia; ?"Germany"
=punctulatus Voet, 1778; spn
=spretus Dejean, 1826 NW-Croatia, NW-Bosnia; NE-Italy:Trieste env.; "Fiume"(=Rijeka)
~angusticollis Motschulsky, 1859, nec Fabricius, 1801 "Carniolia"
=cordicollis Motschulsky, 1865 "Carniolia"(LT)
=imminutus Kraatz, 1879 Croatia:Pljesevica Mts.
~sublineatus Géhin, 1885; un "Südrussland"(error)
~tauricus Ganglbauer, 1888, nec Bonelli, 1811 "Krym"(error)
=basilicatus Born, 1902 C,N-Italy(Basilicata:Val Rondina)
=pseudorugifer Sokolar, 1906 "Strassnitz"(=SE-Moravia:Strážnice)
=occidentalis Born, 1907 CW-France:Vienne:Poitiers
~pueli Lapouge, 1909; un Bouches-du-Rhone:Ile de la Camargue
~sabinus Lapouge, 1909; un C-Italy:?
=cansiglianus Bernau, 1914 NE-Italy:Veneto:Bosco di Cansiglio
~italicus Lapouge, 1924, nec Dejean, 1826 "Basses Alpes:Riez"(=Alpes-Hte-Provence)
=altercoriaceus Kolbe, 1925 "Neumark"(=Gorzów env.)
=italianus Csiki, 1927; nom.pro italicus Lapouge "Basses Alpes:Riez"(=Alpes-Hte-Provence)
=capellae Depoli, 1929 "Capella"(=Croatia:Kapela Massif)
~gracilis Depoli, 1929, nec Küster, 1846 "Mte.Milonia & Timavo valley"(=?Rijeka env.)
~holoradiatus Raynaud, 1966; un
~alaricus (Meyer, 1990); un
=georgii Thelot, 2004 ?

ssp. banaticus Redtenbacher, 1849 S-Hungary; CW,SW-Romania; Serbia, Bosnia, Maced.; ?"Austria:Leitha Mts."(error); ?Banat
=subrugosus Kraatz, 1877 Banat:Bazias
~weiratherianus Breuning, 1936; un Montenegro:Lebrsnik
=praeillyricus Szél, 1993 SW-Hungary:Baranya:Villányi Mts.:Szársomlyó

ssp. mediterraneus Born, 1906 S-Italy:Calabria(Cimina, Antonimina)

ssp. excavatus Charpentier, 1825, nec Paykull, 1790; nbm Croatia & ?N-Albania:Adriatic Sea coast; "Pyrenees"(error)
~rugosus Dejean, 1826, nec Fabricius, 1775 Dalmatia:Vergorac
=abchasicus Motschulsky, 1850 "Abkhasia"(error)
=nitidior Reitter, 1881 Dalmatia:"Zara, Lesina"(=Zadar, Hvar Island)
=dalmaticus Géhin, 1885 "Dalmatia"
=proximus Reitter, 1885 Bosna & Hercegovina:?
=monastiriensis Lapouge, 1922

ssp. florinensis Lapouge, 1922 S-Macedonia; N,C-Greece:Makedonía("Florina:Camp.Grosetti"=Florina env.), Pindos Mt.R.
~pindicola (Maøan, 1952); un N-Pindos Mts.:Malakassion(alpine form)

ssp. ~joniensis Breuning, 1937; un; nom.pro graecus Dejean S-Albania; NW-Greece:Ioannina env; Kerkyra Island; Ionian Isles
~graecus Dejean, 1829, nec Dejean, 1826 "Ionian Isles"
~nitens Gautier des Cottes, 1866, nec Linné, 1758 "Ionian Isles"
=albanicus Breuning, 1943 "S-Albania:Maj Tartarit"
~hickerianus Mandl, 1964; un "Zante Island:Limne Makry"(=Zakynthos)
=zanteanus Breuning, 1964 "Zante Island"(=Zakynthos)

ssp. cerisyi Dejean, 1826 CW,SW-Turkey:Izmir, Manisa, Antalya, Konya; S-Aegean Isles(Mytilene=Lesvos)
=?vicinus Waltl, 1838 "Balkan, Constantinopel"(=Istanbul; ?inexact)
~punctulatus Reiche & Saulcy, 1855, nec Schaller, 1783 Syra(=Syros Island)
~melancholicus Gistl, 1857, nec Fabricius, 1798 "Smyrna"(=Izmir)
=impudicus Gautier des Cottes, 1866 Rhodos
~minor Gautier des Cottes, 1866, nec Kraatz, 1860 "Melos Island"(=Milos)
~sphodrinus Gautier des Cottes, 1866, nec Fischer, 1844 Rhodos
=olympicus Géhin, 1885 "Bithynian Olymp"(=Bursa:Ulu Dag Massif)
~icaricus Ganglbauer, 1888;un Samos & Ikaria Island
~brussensis Lapouge, 1909; un "Brussa"(=Bursa)
=bodemeyeri Lapouge, 1924 ?Bilecik, Karakuey; ?Aksehir, Konya
~kykladicus Breuning, 1936; un; nom.pro punctulatus Reiche & Saulcy Syra(=Syros Island)
~bodemeyerianus (Maøan, 1952); un; nom.pro bodemeyeri Lapouge ?Bilecik, Karakuey; ?Aksehir, Konya

ssp. foudrasi Dejean, 1829 S-Greece:Peloponnesos; Kythera Isl.; Morea(=Peloponnesos)
=obtritus Chaudoir, 1857 "Graecia"
~deplanatus Motschulsky, 1859, nec Fischer, 1823 "Graecia"
~coraxensis Beheim & Breuning, 1943; un S-Greece:"Korax Mts."

ssp. mopsucrenae Peyron, 1858 SC-Turkey:Içel:Tarsus, Mersin; Adana

ssp. hopffgarteni Kraatz, 1878 E-Greece, SW,C-Bulgaria, SE-Serbia(Mt.Komska), E-Macedonia
~semipunctatus Géhin, 1885, nec Donovan, 1806 Rila Mt.R.
=emgei Ganglbauer, 1888 NE-Greece:Thessaloniki, Olympos & Ossa Massif, Parnassos Mt.R.; Euvoia Island
~oetensis (Maøan, 1952); un CE-Greece:Oeta(=?Oite Massif NW of Parnassos)
~olympicola (Maøan, 1952); un NE-Greece:Olympos Massif, 2600m(alpine form)
~storkani (Maøan, 1952); un NE-Greece:Alexandroupoli
~xanthiensis (Maøan, 1952); un NE-Greece:Xanthi(=Xanthé)
~borchmanni Mandl, 1959; un "Stehni-Euböa"(=Euvoia Island)
=deorum (Jeanne, 1970) NE-Greece:Olympos Massif:W-slope:Mt.Agios-Antonios, 2000m
=roeri Blumenthal, 1976 NE-Greece:Chalkidiki:Vaydos

ssp. ressli Mandl, 1961 SE-Turkey:Antakya:Iskenderun env.; betw.Malatya/Elazig; NW-Syria
~montisgavurensis Mandl, 1975; un SE-Turkey:Adana/Gaziantep:Nurdagi Pass

ssp. kindermanni Waltl, 1838 SE-Bulgaria:Burgas, Sliven env.; NW-Turkey; ?SE-Romania; "Konstantinopel"(=Istanbul)
=caraboides Waltl, 1838 "Balkan"
=dobrudjensis Born, 1902 SE-Romania:Mangalia
~pseudofurax (Maøan, 1952); un "Kalofer Balkan"(=Stara Planina Mt.R.nr.Kalofer)
~mehmeti Battoni & Blumenthal, 1973; un Adapazari:Gök Dag nr.Sapanca

ssp. rugifer Kraatz, 1877 Carpathians(?C-Romania:Sinaia)
~rugulosus Ganglbauer, 1888; un C-Romania:Azuga
~testis Lapouge, 1909; un C-Romania:Sinaia

ssp. furax Csiki, 1927; nom.pro montandoni Born SC-Romania(Vlasca:Comana)
~montandoni Born, 1902, nec Buysson, 1882 Vlasca:Comana



6.impressus Klug, 1832, nec Panzer, 1797; nbm SE-Turkey, Syria, Lebanon, Israel, Jordan; C-Iraq:Baghdad env.; ?Sinai Peninsula

ssp. impressus Klug, 1832, nec Panzer, 1797; nbm SE-Turkey:Antakya env.; Lebanon, CW-Syria; C-Iraq:Habbanyiah Lake W of Baghdad
~anxius Lapouge, 1914, nec Duftschmid, 1812

ssp. hybridus Ganglbauer, 1887 SW-Lebanon & W-Israel:sea coast; ?NE-Egypt:Sinai:El Arish; Syria(=currently Lebanon)
~saronicus Lapouge, 1907; un "Jaffa"

ssp. carmelita Lapouge, 1907 N,C-Israel("Galilea, Mt.Carmel, Nazareth")

ssp. palaestinus Lapouge, 1907; ?un C-Israel:N,W of Yerushalaim("Jerusalem"; LT with no locality data); Antilebanon Mt.R.

ssp. dani Schweiger, 1970 NE-Israel:Jordan River source area(Banyas), ?Hermon Massif

ssp. hureishanus Schweiger, 1970 SW-Israel:Negev region:"Djebel Hureisha"(=?Har Ramon Mts.); ?NE-Egypt:C-Sinai

ssp. impressulus Schweiger, 1970 NW-Israel:Haifa env., Elon; SW-Lebanon:Saida env.

ssp. jordanicus Schweiger, 1970 CE-Israel:E of Yerushalaim(Wadi Makkuk/Rijan NW of Jericho)

ssp. negevensis Schweiger, 1970 S-Israel:Negev region, Beer Sheva env.

ssp. simulacrum Rapuzzi, 1997 SE-Turkey:Gaziantep(ca.10-15km W Kilis); N-Syria:Cyrrhus(=W of Kilis)

ssp. inflatior Deuve, 1998 NW-Syria:"Alep env."(=Haleb); 4f

ssp. foveolatior Deuve, 1998 N-Syria:"Jesr’al"

ssp. bourgoini Deuve, 1998 W-Jordan:Petra; 1m

ssp. halfeticus Korell, 1999 SE-Turkey:Sanli Urfa:Halfeti

ssp. palmyrus Görgner, 1999 C-Syria:Tamdoria:97km NW Palmyra:22km SE Uqayribad

ssp. desertum Rapuzzi & Kleinfeld, 2000 SW-Syria:Dimashq env.("W-Damascus:Antilibano, 5-12km NE Assad vill., 1000m")

ssp. drusum Rapuzzi & Kleinfeld, 2000 SW-Syria:Jebel al Druz Mts.:Es Suweida, 1050-1600m

ssp. neojordanicus Kleinfeld & Rapuzzi, 2004 NW-Jordan:Amman env.(Wadi Sir near Amman)



7.mulsantianus Morawitz, 1886; nom.pro asperatus Mulsant SC,SE-Turkey:Mersin to Antakya; NW-Syria

ssp. mulsantianus Morawitz, 1886; nom.pro asperatus Mulsant SC-Turkey:Mersin:Gülek; Adana; Antakya; ?"Cilicia:Adana env."
~asperatus Mulsant, 1853, nec (Dejean, 1826) ?"Cilicia:Adana env."
=kotschyi Ganglbauer, 1887 ?"Cilician Taurus"
~akbesianus Lapouge, 1914; un Antakya:"Akbes"(nr.Iskenderun)
=aleppensis Lapouge, 1914 NW-Syria:Haleb env.("Aleppo"=Haleb)
~adspersarius Lapouge, 1917; un ?"Cilicia"
~nurdaghensis Blumenthal, 1967; un Antakya:Nur Dagi:Kirikhan env.

ssp. bernhauerorum Heinz, 1977 SE-Turkey:Kahramanmaras:pass S of Göksun



8.payafa White, 1845 SC-Turkey

ssp. payafa White, 1845 S-Turkey:Mugla, Antalya, Isparta; "Lycia:Günek"
=pseudoacuticollis Èatloš & Moravec, 1999 SW-Turkey:Antalya:Bademli Pass

ssp. acuticollis Motschulsky, 1858 SC-Turkey:Antalya, Icel, Adana, Kahramanmaras; "Kurdistan"(inexact); alpine form
=drouxianus Machard, 1986 Konya:N of Ermenek:Yellibeli Pass(alpine form)



9.piochardi Géhin, 1883 Lebanon, N-Israel, NW-Syria, NE-Turkey:Antakya

ssp. piochardi Géhin, 1883 C-Lebanon:E of Beyrut:Djebel Sannin(LT)

ssp. praestigiator Morawitz, 1886; nom.pro ehrenbergi Klug, 1832 N,C-Lebanon; "Syria"(currently Lebanon)
~ehrenbergi Klug, 1832, nec Fischer, 1829 "Syria"(currently Lebanon)

ssp. azari Deuve, 1998 N-Lebanon:Fnaideq env.:Mts.W of Mechmech, 1000m

ssp. pierpaolo Kleinfeld & Rapuzzi, 2004 N-Lebanon:Bcharré env. (El Arz, 1900m)

ssp. labruleriei Géhin, 1883 S-Lebanon:Dj.Cheik, El Hebbariyé, Ebel es Saqi, Marjaayoun; N-Israel:Zefat, Elon env.
=bytinskii Schweiger, 1970 N-Israel:Elon

ssp. bardawilae Deuve, 2002 S-Lebanon:Rachaya

> ssp. morawitzi Ganglbauer, 1887 SE-Turkey:Antakya, Gaziantep; NW-Syria:Al Ladhiqiyah env.; "Asia Minor"
=montesamanus Mandl, 1967 SW-Turkey:Gaziantep:Amanus Mts.:Sakçagözü:Fevzipaºa env.
=discipulus Rapuzzi, 1997 Antakya:betw.Reyhanli/Cilvegozu
=tainei Lassalle, 1998 Antakya:Yayla Dag, 800m
=grososae Deuve & Remond, 1998 NW-Syria:Halab env.

ssp. gezei Deuve, 2002 N-Lebanon(Qammouaa)

ssp. pinguis Lapouge, 1914 "Palestine:Nazareth"(=N-Israel:Nazerat; NT)



10.talyschensis Ménétriés, 1832 Azerbaydzhan:Talysh; N-Iran:Gilan & Elborz Mt.R.

ssp. talyschensis Ménétriés, 1832 Azerbaydzhan:Talysh; N-Iran:Gilan Mt.R.; W,C-Elborz Mt.R.
~elongatus Motschulsky, 1859, nec Duftschmid, 1812 "conf.Pers."
~glabripennis Mandl, 1953; un "Mazenderan"

ssp. luctuosior Deuve, 1994; nom.pro luctuosus Zoubkoff NE-Iran:E-Elborz Mt.R.:"Astrabad"(=Gorgan); SW-Turkmenistan:Kopetdag
~luctuosus Zoubkoff, 1837, nec Villers, 1789 "Astrabad"(=Gorgan)


BACK



LAMPROSTUS Motschulsky, 1865; type species Carabus spinolae Cristoforis & Jan = Carabus torosus spinolae Cristoforis & Jan
species: 12



1.calleyi Fischer, 1823 Georgia, Armenia, E-Turkey; Azerbaydzhan:Talysh env.; N-Iran:Gilan Mt.R.

ssp. calleyi Fischer, 1823 Georgia, E-Caucasus; "Transcaucasia:Georgia/Persia borders"

ssp. prasinus Ménétriés, 1832, nec Thunberg, 1784; nbm N-Iran:Gilan Mt.R.; Azerbaydzhan:Lenkoran env.; "Talysh:Perimbal nr.Lenkoran"; LT
=prasinescens Deuve, 1994; nom.pro prasinus Ménétriés

ssp. prevosti Gory, 1833 Georgia:?Akhaltsikhe env.; "Sibiria"(by error; LT)
=renardi Chaudoir, 1846Georgia:"Abastumani nr.Achalzich"(=Akhaltsikhe; LT)
=thermarum Motschulsky, 1850 Georgia:"Achalzik"(=Akhaltsikhe; LT)
~procrustoides Lapouge, 1909, nec Géhin, 1885

ssp. pseudoprasinus Lapouge, 1912 S-Armenia; S-Georgia; "Transcaucasia"
=manderstjerni Motschulsky, 1852 "Alp.Cauc., Persia"(LT)
~pseudocalleyi Lapouge, 1914; un; error for pseudoprasinus Lap., 1912
=nigropolitulus Mandl, 1970 Armenia:?Sevan Lake
=araxenus Jablokov-Khnzorjan, 1976 ?Araks valley

ssp. subnigrinus Heinz, 1970 CE-Turkey:Agri:"Tahir Pass"(=Saç Pass betw.Tahir/Horasan)

ssp. martinae Machard, 1977 NE-Turkey:Artvin:Yalnizcam Pass; Kars:Kars env., Göle, Sarikamis
=catakensis Machard, 1985 Kars:Catak nr.Sarikamis

ssp. chaudoiriellus Deuve, 1995 "Caucasus"; ?Dagestan:Bogosskiy Mt.R.

ssp. chalconatus Mannerheim, 1830 E-Turkey:Gümüshane, Bayburt(Kop Dag), Erzurum, Sivas, Adana, Kirsehir
=luxuriosus Motschulsky, 1844
=chalcochlorus Chaudoir, 1852 SE-Turkey:Diyarbakir(probably by error)
~ambiguus Lapouge, 1914, nec Paykull, 1790
=hozari (Maøan, 1952) Kirºehir:Mucur
~tschimenensis Breuning, 1964; un Tschimen Dag nr.Bayburt
=hisarensis Darge, 1989; non junior homonym Sivas/Ordu:old road betw.Koyulhisar/Mesudiye
=koyunensis Darge, 1990; nom.pro hisarensis Darge Sivas/Ordu:old road betw.Koyulhisar/Mesudiye
=maguini Auvray, 1994 Sivas:Corakbogazi Pass S of Imranli

ssp. subnigritulus Heinz, 1990 E-Turkey:Erzurum:W of Narman:Kirecli Pass

ssp. claudeauvrayi Cavazzuti, 2001 E-Turkey:Gümüshane:Kostandagi Pass, 2600-2700m

ssp. nigrinus Motschulsky, 1865 Azerbaydzhan:Talysh env.; N-Iran:Gilan Mt.R.; Conf.Pers.Lenkoran(LT)
= rechingerae Mandl, 1972 N-Iran:Azarbaijan:Dasht Moghan:betw.Ardabil/Parsabad(=near Guermi); 1f

ssp. inseriatus Mandl, 1955 N-Iran:Azarbaijan:W of Ardabil:Nir env.; "Persia"
=nirensis Heinz, 1970 N-Iran:Azarbaijan:W of Ardabil:Nir

ssp. machardiellus Deuve, 1991; nom.pro kurdistanicus Machard CE-Turkey:Van:Güzeldere Pass nr.Baskale
~kurdistanicus Machard, 1986, nec Eidam, 1931 Van:Güzeldere Pass nr.Baskale



2.cylindricus Lapouge, 1914, nec Herbst, 1784; nbm N-Iran:Elborz Mt.R.(probably Rasht env.; NT)
=cylindricior Deuve, 1994; nom.pro cylindricus Lapouge
=staveni Heinz, 2002 N-Iran:Gilan:S of Dailiman (Siah-Sar); Durfak env.



3.erenleriensis Schweiger, 1966 NW-Turkey:Bolu to Cankiri

ssp. erenleriensis Schweiger, 1966 NW-Turkey:Bolu:betw.Bolu/Mudurnu(Abant Mts.); Kaynasli env.
~erenleriensis Schweiger, 1962; un Bolu:Abant Mts.
=kaynasliensis Machard, 1975 Bolu:betw.Bolu/Duzce:Kaynasli

ssp. bithynicus Heinz & Korge, 1967 NW-Turkey:Bolu:betw.Duzce/Akcakoca(S of Akcakoca)

ssp. pseudotorosus Schweiger, 1969 NW-Turkey:Bolu:SW of Mengen

ssp. fortepunctatus Battoni & Blum., 1973; nom.pro rotundicollis Blum.& Br. NW-Turkey:Cankiri:Eskipazar; NE of Bolu:Mengen, Yedigöller ~rotundicollis Blumenthal & Breuning, 1966; un NE of Bolu:Mengen

ssp. karadaghensis Battoni & Blumenthal, 1973 NW-Turkey:Adapazari:Kara Dag Massif S of Akyazi, Dogancay env.

ssp. ahmetustaensis Cavazzuti, 1987 NW-Turkey:Zonguldak:Ahmetusta Pass, Eflani

ssp. hamus Basquin & Legrand, 1991 NC-Turkey:Ankara:Camlidere



4.guycolasianus Deuve, 2001 N-Iran:Gilan, Mazandaran

ssp. guycolasianus Deuve, 2001 N-Iran:Mazandaran:100km S Chalus; Gilan:Rostamabad

ssp. colorescens Deuve, 2001 N-Iran:Elborz Mt.R:?("Elburs")



5.hemprichi Dejean, 1826 Lebanon, W-Syria, Israel, NE-Turkey:Antakya; ?Sinai Peninsula

ssp. hemprichi Dejean, 1826 N,C-Lebanon, W-Syria:Homs, Haleb; "Syria"
~libanicus Lapouge, 1907; un "Lebanon"
~spaethi Lapouge, 1907; un LNF
~ruginus Lapouge, 1914; un

ssp. damascenus Lapouge, 1924 SW-Syria:Dimashq env.("Damascus"), Baalbek; NE-Israel:Jordan River sources

ssp. propheta Rapuzzi, 1995 SE-Turkey:Antakya:betw.Reyhanli/Cilvegozu

ssp. sidonius Lapouge, 1907; ?un SW-Lebanon(Saida) & W-Israel:sea coast; ?NE-Egypt:Sinai:El Arish
~epistomalis Lapouge, 1907; un Saida

ssp. elonensis Schweiger, 1970 C,N-Israel(Elon)

ssp. cheikhermonensis Deuve, 1992 Lebanon:Antilebanon Mt.R.:E of Saida:Djebel-ech-Cheik



6.ledouxi Deuve, 2001 N-Iran:Gilan/Azarbaijan:Khalkhal(E of Heroabad:Mt.Charasu Dagh), Ardabil, Asalam
> ~pseudonirensis Blumenthal, in litt.



7.punctatus Castelnau, 1835 NW-Syria; N-Lebanon; NE-Turkey:Antakya; "Syria"
~duponcheli Barthelemy, 1837, nec Dejean, 1831
=durvillei Dupont, 1838
=judaicus Géhin, 1885 "Syria:Jbel Sannin"(=C-Lebanon:E of Beyrut:Djebel Sannin)
=cristatopunctatus Lapouge, 1907
~vagepunctatus Lapouge, 1907; un N-Lebanon
=koubbii Deuve, 1998 N-Syria:"Jesr’al"



8.robustus Deyrolle, 1869 NE-Turkey:Giresun to Rize; ?Georgia:Batumi

ssp. robustus Deyrolle, 1869 NE-Turkey:Trabzon env.; ?Georgia:Batumi; "Trébizonde"(=Trabzon; LT)
=callipeplus Schaufuss, 1882
=extremus Schaufuss, 1882
=perfectus Schaufuss, 1882
=semirugosus Schaufuss, 1882
~sexpunctatus Schaufuss, 1882, nec Linné, 1758
=triumphator Schaufuss, 1882
=colossus Bleuse, 1896
=trapesunticus Eichler, 1924 "Elleon nr.Trapezunt"(=Trabzon)

ssp. ducalis Schaufuss, 1882 NE-Turkey:Trabzon:Macka, Zigana Pass; Rize:Kaptanpasa; LNF
~singanensis Breuning, 1932; un NE-Turkey:Trabzon:"Singana Dag"(=Mts.nr.Zigana Pass)
=subrobustus Machard, 1994 Rize:Dereköy SE of Ikizdere

ssp. athleta Semenov, 1896 NE-Turkey:Gümüshane:Köse, Siran, Kelkit env.

ssp. triumphans Cavazzuti, 1987 NE-Turkey:Giresun:Tirebolu env.(Eymur Köyü, 150m); Trabzon:Tonya env.
=exsultans Cavazzuti, 1993 NE-Turkey:Giresun:Espiye env.

ssp. kaptanpasa Cavazzuti, 2001 E-Turkey:Rize:Kaptanpasa, 1500m

ssp. pseudorobustus Blumenthal & Heinz, 1977 NE-Turkey:Giresun:Kulakkaya(exactly: ca.10km S Yavuzkemal); Kümbet env.
=yavuzkemalensis Deuve & Remond, 2001 NE-Turkey:Giresun:Yavuzkemal, 1300m; btw. Pinarlar/Yavuzkemal, 1000m
=pseudocolossus Cavazzuti, 1993 NE-Turkey:Giresun:betw.Giresun/Dereli:Konacik

ssp. egribelensis Deuve & Remond, 2001 NE-Turkey:Giresun:N of Egribel Pass; 1600-1700m



9.rostandianus Deuve, 2005 "Lebanon:Farm, May 65"; 1m



10.saulcyi Piochard, 1875 N-Lebanon, NW-Syria

ssp. saulcyi Piochard, 1875 N-Lebanon:Lebanon Mt.R.:Eden(=Ehden),"Khamés"(LT)
=chambouqensis Lassalle, 1998 N-Lebanon:Chambouq

ssp. jitkae Deuve & Macek, 1990 NW-Syria:Al Ladhiqiyah:Djebel Ansariya Mts.(Qerdaha env.)

ssp. ehdenensis Deuve, 1992 N-Lebanon:Ehden(?by error; 1m)



11.syrus Roeschke, 1898 Lebanon

ssp. syrus Roeschke, 1898 Lebanon:Lebanon Mt.R.:E of Beyrut:Djebel Sannin

ssp. cheikensis Deuve, 1992 Lebanon:Antilebanon Mt.R.:Djebel-ech-Cheik

ssp. saidaensis Deuve, 1992 S-Lebanon:Saida; 1m

ssp. pseudopinguis Heinz & Staven, 2000 N-Jordan:NW of Jarash:Sakhra, Ajlun; W of Jarash:SW of Anjarah

ssp. moukarzelae Deuve, 2002 S-Lebanon:Marjaayoun

ssp. eidae Deuve, 2002 S-Lebanon:Rachaya

ssp. lecordieri Deuve, 1992 "Upper Jordan:Khent"; 1m



12.torosus Frivaldsky, 1835 E-Bulgaria; NW,C,NE-Turkey; ?SE-Romania

ssp. torosus Frivaldsky, 1835 E-Bulgaria; ?SE-Romania; "E-Balcans:Black Sea coast"

ssp. jani Géhin, 1885; nom.pro moestus Cristoforis & Jan, 1812 CW-Turkey:Bilecik, Eskisehir, Konya, Afyon, Kayseri, Adana, Icel, Isparta
~moestus Cristoforis & Jan, 1837, nec Duftschmid, 1812 Eskisehir
~escherichi Lapouge, 1905; un Konya:Sultandag nr.Aksehir; Afyon:Emirdag
~bodemeyeri Lapouge, 1914; un Konya
~mundus Lapouge, 1914; un "Damascus"(error)
=hoberlandti (Maøan, 1952) Erciyas Dag, Bolkar Dag, Bakir Dag
=gajaci Breuning, 1979 Isparta:Davraz Dag
=hahni Mandl, 1980 Konya:Karapinar env.

ssp. procrustoides Géhin, 1885 C-Turkey:Diyarbakir, Malatya

ssp. pseudescherichi Breuning, 1962 C-Turkey:Konya:Tuz Lake:E-coast
=vignatagliantii Breuning & Ruspoli, 1977 Konya:Tuz Lake env.

ssp. cankirianus Blumenthal & Breuning, 1967 NW-Turkey:Cankiri:betw.Cankiri/Ilgaz(steppes N of Cankiri), Cerkes env.; Ankara:Güvem
=basquinianus Darge, 1992 Ankara:betw.Yukaricanli/Güvem
=crudus Darge, 1992 Cankiri:Sacakbeli Pass
=xanthicus Darge, 1992 Cankiri:Kurtcimeni env.

=reynieri Charet, 1992 Cankiri:Sacakbeli Pass betw.Cerkes/Sabanözü

ssp. leonidas Cavazzuti, 1999 SW-Turkey:Isparta:Gelendost

ssp. spinolae Cristoforis & Jan, 1837; ?non priority C-Turkey:Amasya(NT), Samsun, Ankara, Kirsehir, Yozgat, Corum, Tokat
=bonplandi Ménétriés, 1837; ?priority "environs od Constantinople"(=Istanbul; ?inexact)
=lamprus Chaudoir, 1850 Samsun; "Anatolia"
=phaedimus Schaufuss, 1882 Amasya
~obesus Lapouge, 1909; un Tokat
=mercatii Breuning & Ruspoli, 1970 Tokat:Mamo Dag nr.Almus(=ca.10km WNW Almus)

ssp. salignus Schweiger, 1969 NC-Turkey:Samsun:Kavak

ssp. vexator Schweiger, 1969 NC-Turkey:Amasya:Merzifon; Samsun:Ladik env.

ssp. ladikensis Deuve & Charet, 1992 NC-Turkey:Samsun:Ladik

ssp. nordmanni Chaudoir, 1848 NE-Turkey:Black Sea coast betw.Samsun/Ordu; LT without locality data
=productus Hampe, 1852 LT without locality data
=rabaroni Breuning, 1964 Ordu:forest zone nr. Akkus
=enricoi Battoni & Blumenthal, 1973 Ordu:betw.Gölköy/Mesudiye
=sturanicus Breuning & Ruspoli, 1979 Samsun:Akpinar nr.Carsamba
=dominici Thomé, 1987 Ordu:ca.5km S Mesudiye

ssp. giresuni Breuning, 1964 NE-Turkey:forest zone above Giresun/Ordu; betw. Dereli/Yavuzkemal; Egribel & Sehitler Pass
~paululus Heinz & Korge, 1964; un Giresun:Egribel Pass(alpine form)
=drouxi Machard, 1977 NE-Turkey:Giresun:betw.Dereli/Kümbet, 1500m

ssp. pseudoducalis Heinz & Korge, 1967 NW-Turkey:Kastamonu:betw.Ilgaz/Kastamonu(Ilgaz Dag Massif)
~pseudoducalis Schweiger, 1964; un Kastamonu:Ilgaz Dag Massif

ssp. usurpator Cavazzuti, 1987 NW-Turkey:Kastamonu:Devrekani env.(N of Devrekani)

ssp. sinopensis Breuning & Ruspoli, 1970 NC-Turkey:Sinop:Dranaz Dag
~sinopensis Breuning, 1961; un Sinop:Dranaz Dag
~caeruleoviolaceus Breuning & Ruspoli, 1970; un betw.Dranaz Dag/Sinop

ssp. lohsei Blumenthal & Heinz, 1978 NC-Turkey:Kastamonu & Sinop:Devrekani to Sinop(Yaraligöz Massif)

ssp. rouziei Breuning & Ruspoli, 1970 NC-Turkey:Tokat(Dumanli Dag nr.Almus=SE of Almus), Sivas

ssp. rigouti Basquin & Darge, 1986 NC-Turkey:Sivas:Koyulhisar env.


BACK



MACROTHORAX Desmarest, 1850; type species Carabus aumonti Lucas
species: 4



1.aumonti Lucas, 1849 NW-Algeria, N-Morocco

ssp. aumonti Lucas, 1849 NW-Algeria:"Djemaa nr.Nemours"(=Ghazaouet), Maghnia; NE-Morocco:Oujda env.
=lepitrei Krausse, 1910 Morocco:Angad plateau(=betw.Oujda/Maghnia)
=sculptus Lapouge, 1925 NW-Algeria:"Lalla Maghrnia"(=Maghnia)

ssp. maroccanus Bedel, 1895 N-Morocco:Rif Mts., Melilla, Taza, Sefrou, Meknes
=lapougei Krausse, 1910
~riffensis Boettger, 1921; un N-Morocco:Rif Mts.:?



2.morbillosus Fabricius, 1792 NW-Africa, S-Spain, Baleares, Sicily, Sardinia, Corsica, S-Italy:Calabria

ssp. morbillosus Fabricius, 1792 NC,NE-Algeria:Alger env.:Medea, Blida, Tizi Ouzou, Mt.Djurdjura; Bejaia; "Barbaria"
=mittrei Lucas, 1849 NE-Algeria:"Bougie env."(=Bejaia)
=cheminorum Deuve, 1988 S-France:Var:Frejus; introduced

ssp. alternans Palliardi, 1825, nec Fabricius, 1792; nbm Sicily; S-Italy:Calabria
=servillei Solier, 1835
~viridulus Ragusa, 1905, nec Kraatz, 1878
=bruttianus Born, 1906 Calabria:Aspromonte massif:Sta.Eufemia d'Aspromonte
~borni Krausse, 1908, nec Meyer-Darcis, 1902
=sicanus Csiki, 1927; nom.pro borni Krausse

ssp. constantinus Kraatz, 1899 NE-Algeria:Constantine, Setif; N,CE-Tunisia; NW-Libya:Yafran; Sardinia, Corsica, Lampedusa
=arborensis Krausse, 1908 Sardinia:Asuni
=galloprovincialis Lapouge, 1910 S-France:Var:Muy; introduced
~relictus Krausse, 1910, nec Semenov, 1888 S-France:Les Maures; introduced
=corsicanus Lapouge, 1913 Corsica
=lampedusae Born, 1925 Lampedusa Island
~bodemeyeri Kolbe, 1928, nec Lapouge, 1924 Tunisia:Ain-Draham nr.Tebarca(=Tebarka)
=mirei Breuning, 1975 NE-Algeria:Setif:Djebel Babor Massif

ssp. cychrisans Lapouge, 1899 NW-Algeria:Oran env. ?Maghnia
~marginatus Lallemant, 1868, nec Linné, 1758 Oran
=oranicus Csiki, 1927; nom.pro marginatus Lallemant NW-Algeria:Oran

ssp. macilentus Lapouge, 1899, nec Olivier, 1795; nbm S-Spain:Alicante, Murcia; Baleares
=ferrugatus Lapouge, 1899 Baleares
=balearicus Lapouge, 1913 Baleares



3.planatus Chaudoir, 1843, nec Duftschmid, 1812; nbm N-Sicily; LNF(unlabelled NT)
=thomsoni Fairmaire, 1857



4.rugosus Fabricius, 1792 N,C-Morocco, Spain, Portugal; introduced:SC-France

ssp. rugosus Fabricius, 1792 N,C-Morocco:Oued Tensift-Marrakech-Taza-Tanger-Ribat-Casablanca
=barbarus Dejean, 1826 "Africa"(?Tanger)
=macurebus Géhin, 1885 "Morocco:Tlemcen"(=Algeria:W of Oran; ?error)
=richteri Quedenfeldt, 1887 Morocco:Casablanca
~tingitanus Lapouge, 1902; un Morocco:?Tanger
=beidensis Codina, 1927 Morocco:Casablanca
=promeces Alluaud, 1930 Morocco:Oued Cherrat(=betw.Dar el Beida/Ribat:nr.Ben Slimane)
=pseudorichteri M.Machard & P.Machard, 1993 Morocco:SW of Afourer:El Tleta-Ouzoud; 1m

ssp. celtibericus Germar, 1824 Portugal; W-Spain:Galicia - Caceres - Huelva; Portugal
=brannani Schaufuss, 1871 S-Portugal:?Portalegre; ?Sierra de Estrella
~semicyanescens Bra?es, 1985; un
=pseudoboeticus Lassalle, 1986 SW-Spain:Huelva:La Palma del Condado
=condadoensis Bra?es, 1992; nom.pro pseudoboeticus Lassalle SW-Spain:Huelva:La Palma del Condado

ssp. laufferi Breuning, 1927 C-Spain:Avila env.

ssp. baeticus Deyrolle, 1852 S-Spain:Sevilla - Granada - Almeria; "Cordilléres Bétiques"
~andalusiacus Géhin, 1885; un Andalusia
=seguranus Lauffer, 1905 Jaen/Granada:Sierra de Segura Mts.(alpine form)
=breuili Alluaud, 1930 Andalusia:Laguna de Janda
~pseudobaeticus Bra?es, 1985; un Granada:?

ssp. levantinus Lauffer, 1905 E-Spain(Murcia & Valencia); SC-France:Pyrenées-Orientales(?introduced)
=barnolai Xaxars, 1921 Barcelona:St.Boy de Llobregat
=vermiculosus Fons, Leplat & Ferrero, 1993 SC-France:Pyrenées-Orientales:Albéres Massif:E-slope; ?introduced



RELICTOCARABUS Ledoux, 1984; type species Relictocarabus meurguesae Ledoux = Carabus meurguesianus Ledoux
species: 1



1.meurguesianus Ledoux, 1990; nom. meurguesae (Ledoux)Morocco:High Atlas:Djebel Toubkal Massif:Oukaimeden env.; "Grand Atlas"
~meurguesae (Ledoux, 1984), nec Ledoux, 1980 "Grand Atlas"


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APOPLESIUS Deuve, 1990; type species Carabus omphreodes Reitter
species: 4



1.deuvei Lassalle, 1986 CE-Turkey:Tunceli, Muº

ssp. deuvei Lassalle, 1986 CE-Turkey:Tunceli:Pülümür, 1600m

ssp. hajdaji Häckel, 2007 CE-Turkey:Muº:60km NW Muº:Bulgan Geçidi, 38°56'N/41°08'E, 1630 m; 1m



2.legrandi Deuve, 1991 NE-Turkey:Kars:Digor



3.omphreodes Reitter, 1898 SE-Turkey:Mardin; 1m



4.pseudodepressus Machard, 1988 SE-Turkey:Bitlis:S-coast of Van Lake:Resadiye; 1m/1f


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INIOPACHUS Solier, 1848; type species Carabus pyrenaeus Audinet-Serville
species: 2



1.auriculatus Putzeys, 1872 NW-Spain:Cantabrian Mts.

ssp. auriculatus Putzeys, 1872 NW-Spain:Cantabrian Mts.:Picos de Europa Massif
=impensus Reitter, 1896Picos de Europa Massif
~emmanueli Prunier, 1993; un

ssp. ubinensis (Puisségur, 1955) NW-Spain:Cantabrian Mts.:Pena Ubina Massif
~mestrei (Puisségur, 1955); un
=ornizensis Meyer, 1998 Cantabrian Mts.:Pena de Orniz



2.pyrenaeus Audinet-Serville, 1821 N-Spain & S-France:Pyrenees

ssp. pyrenaeus Audinet-Serville, 1821 W-Pyrenees:Pic d’Orhy to Pic de Vignemale Massif; Hautes Pyrenées
~punctatus Géhin, 1885, nec Castelnau, 1835
=ignitus Reitter, 1896 Hautes Pyrenées
~cephalotes Nicolas, 1919, nec Linné, 1758 Pyrenees-Atlantiques:Sede-de-Pan
=cephalicus Csiki, 1927; nom.pro cephalotes Nicolas Pyrenees-Atlantiques:Sede-de-Pan
~punctatulus Csiki, 1927; un; nom.pro punctatus Géhin
~punctatulus (Jeannel, 1941), nec Fabricius, 1792
~occidentalis Jeanne, 1969, nec Born, 1907 Pyrénées-Atlantiques:Pic d’Orhy, 2000m

ssp. costulus Géhin, 1885; nom.pro costatus Kraatz Pyrenees-Orientales:Mt.Canigou Massif; "E-Pyrenees"
~costatus Kraatz, 1878, nec Germar, 1824 E-Pyrenees

ssp. cerdanus Lapouge, 1924 Andorra:Mt.Casamanya to Pyr.-Or.:Puigmal Massif; N-Spain:Maladetta nr.Olot:Col d’Envalira
~laurentii Rouselle, 1976; un

ssp. vallierensis Colas, 1965 C-Pyrenees:Ariege, Val d’Aran, Mt.Vallier:Port d’Aula

ssp. jeanneli Deuve, 1994 C-Pyrenees:Pic de Vignemale Massif to Viella env.; Huesca:Pt.de Venasque
~immetallicus Nicolas, 1898; un
~gavarniensis Pater, 1938; un
~viridissimus Pater, 1938; un


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CHRYSOTRIBAX Reitter, 1896; type species Carabus hispanus Fabricius
species: 2



1.hispanus Fabricius, 1787 C,CW,CE,SC-France; "Spain"(error; perhaps: Lozére:Ispaniac)
~hispanicus Thomson, 1875; un; error for hispanus Fabricius
~gabalicus Géhin, 1885; un Lozére
~cevennicus Jacquet, 1886; un SE Massif Central:Cevennes Mts.
=boudeti Beuthin, 1892 Drôme:Foret de Saou
cebennicus Vuillefroy, 1892 SE Massif Central:Cevennes Mts.
=languedocianus Vuillefroy, 1892 HT not designated
~castaneus Lapouge, 1902; un Lozére
=latissimus Lapouge, 1910 Aude:Rennes les Bains
=auzati Le Moult, 1912 Dufourt
=infernalis (Lebis, 1958) Tarn:Foret de la Grésigne
~mourguesi (Puisségur, 1962); un Lozére.Saint-Germain-de-Calberte
=berardi (Raynaud, 1967) Loire:Saint-Victor-sur-Loire
~infuscatus (Raynaud, 1967); un Tarn:Foret de la Grésigne
~semivirescens (Raynaud, 1967); un Aveyron:Villecomptal
~alarici (Darnaud, 1983); un Aude:Rennes les Bains
~canui (Darnaud, 1983); un Corréze:Tulle, Argentat, Beynat
~ignivorax (Darnaud, 1983); un Lozére:St.Germain de Calberte
~retaudis (Gorrin, 1991); un Charente-Maritime, nr.Saintes.Bois de Retaud
~geraulenai Lavagne, 2002; un HT not designated



2.rutilans Dejean, 1826 E-Pyrenees; NE-Spain:Barcelona, Lerida

ssp. rutilans Dejean, 1826 E-Pyrenees:Ariege to Pyr.Or.(Prats de Molo); Gerona to Lerida; Barcelona; Tarragona
~aequatus Lapouge, 1902, nec Motschulsky, 1845 Barcelona:Montserrat
~rusticus Lapouge, 1902; un "NE Spain"
~brevicollis Lapouge, 1910, nec Fabricius, 1792 Aude:Ft.de Monthaut nr.Rennes-les-Bains
~viridis Ferrer y Vert, 1910; un Barcelona:Montserrat
~affictus Codina, 1913; un Setcasas
=aragonensis Lapouge, 1913 Barcelona:Sierra de Montseny
=codinae Lapouge, 1913 Setcasas
~montserraticus Csiki, 1917; un; nom.pro aequatus Lapouge Barcelona:Montserrat
~viridicolor Csiki, 1927; un; nom.pro viridis Ferrer y Vert Barcelona:Montserrat
~interseriatus L’Hoste & Meyer, 1973; un HT not designated
~delacrei Tarrier, 1975; un Barcelona:Berga:Nostra Signora de Queralt
~alberensis (Darnaud, 1978); un E-Pyrenees:Amélie-les-Bains
~blanci (Darnaud, 1978); un E-Pyrenees:Prats-de-Mollo
~lecumberryi (Darnaud, 1978); un E-Pyrenees:Prats-de-Mollo
~pseudoopulentus (Darnaud, 1978); un Barcelona:Sierra de Montseny
~boulbeni (Blanc, 1980); un Gerona:Camprodon
~darnaudi (Blanc, 1980); un Barcelona:Sierra de Montseny
~claveriei Lassalle & Pham, 1981; un Barcelona:Sierra de Montseny
=jeannei Lassalle & Pham, 1981 Tarragona:Sierra de la Mussara
~goalesae (Cottour, 1984); un Barcelona:Sierra de Montseny
~lassallei Bra?es, 1985; un Tarragona:Prades
~alberensis Mollard, 1999; un E-Pyrenees:Alberes Mts.:Ft.de Sorrede
~casteilensis Mollard, 1999; un NE-Spain:E-Pyrenees:near Casteils
~frontanyaensis Mollard, 1999; un NE-Spain:E-Pyrenees:N of Borreda
~merangensis Mollard, 1999; un NE-Spain:N-Meranges:W of Puicerda, 1500-1700m
~quillansis Mollard, 1999; un Aude:S of Quillan
~romaricus Mollard, 1999; un probably: E-Pyrenees:N of Borreda
~soredensis Mollard, 1999; un E-Pyrenees:Alberes Mts.:Ft.de Sorrede
~urgelensis Mollard, 1999; un NE-Spain:E-Pyrenees:N Sierra del Cadi
~palensis Mollard, 1999; un NE-Spain:E-Pyrenees:Pall Pass
~occideneissimus Mollard, 1999; un NE-Spain:E-Pyrenees:Boixols Pass

ssp. perignitus Reitter, 1896 Andorra, N-Spain:Lerida; Pyrenees of Spain
~foveipennis Lapouge, 1902; un Andorra
~fontyi (Darnaud, 1980); un Andorra
~basetaensis Mollard, 1999; un NE-Spain:E-Pyrenees:Sierra de la Baseta
~benenensis Mollard, 1999; un NE-Spain:E-Pyrenees:Sierra de Aigues-Tortes, Benes Valley
~capdellensis Mollard, 1999; un NE-Spain:E-Pyrenees:La Torre de Capdella
~juberriensis Mollard, 1999; un NE-Spain:E-Pyrenees:SE of Andorra:near Juberri
~perversensis Mollard, 1999; un NE-Spain:E-Pyrenees:near Sierra de St.Gervas, near Pervers Pass

ssp. opulentus Oberthür, 1898 NE-Spain:C-Pyrenees:Lerida, Huesca(Col de la Muria)
~alinsis Mollard, 1999; un NE-Spain:Sierra de Monteixo:near Alins
~cotiellensis Mollard, 1999; un NE-Spain:"E Cotiella Mt."
~turbonenesis Mollard, 1999; un NE-Spain:Turbon Mts.:Siarra del Ballabriga


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CHRYSOCARABUS Thomson, 1875; type species Carabus auronitens Fabricius
species: 5



1.auronitens Fabricius, 1792 C,W-Europe:Pyrenees to Bohemia; Alps, Carpathians

ssp. auronitens Fabricius, 1792 C,E-France, Germany, Holl., Belg., Switz., N-Austria, Bohemia, W-Poland; "Sweden"(error)
~auratonitens Illiger, 1798; un; error for auronitens Fabricius
=zwickii Heer, 1837 Switzerland:Schaffhausen
~atratus Heer, 1841, nec Duftschmid, 1812
~coerulescens Letzner, 1850; un
~picipes Letzner, 1850; un
~versicolor Letzner, 1850; un
~viridis Letzner, 1850; un
=putzeysi Mors, 1863 Belgium:?
~nigripes Heyden, 1875; un "Domodossola"
~azureoauratus Dalla Torre, 1877; un
~picipes Dalla Torre, 1877; un
~viridiauratus Dalla Torre, 1877; un
=subcatenulatus Westhoff, 1881
=ignifer Haury, 1889 "Mühlhausen env."(=Mulhouse)
=joerini Stierlin, 1890 Zürich env.
=perviridis Reitter, 1896
=auropurpureus Lapouge, 1898 Belgium:?
=cluniensis Born, 1903 Vorarlberg:Hochfreschen
=petzi Sokolar, 1907 S-Austria:?
=brevipennis Lapouge, 1913 ?"Nordsteiermark"
~gervaisi Le Moult, 1913; un Eure:Foret de Lyons
=allevardensis Nicolas, 1919 Isére:Allevard
~letacqui Antoine, 1919; un N-France:?
~borni Lapouge, 1924, nec Meyer-Darcis, 1902 Switzerland:Emmenthal
~charlottei Venet, 1926; un
=foveipennis Lebis, 1926 E-France:?
~pauli Csiki, 1927, nec Kolbe, 1923; nom.pro borni Lapouge Switzerland:Emmenthal
=foveatipennis Csiki, 1927; nom.pro foveipennis Lebis E-France:?
~oberthueri Lebis, 1928; un ?N-France
~aureus Sirguey, 1931, nec DeGeer, 1774 "Normandia"
~coeruleomicans Sirguey, 1931; un "Normandia"
=guedeli Sirguey, 1931
~marginatus Sirguey, 1931; un "Ft.d’Ecouve & Ft.de Perseigne"
=normannensis Sirguey, 1931 "Normandia"
=normanneus Sirguey, 1931 "Normandia"
~purpuratus Sirguey, 1931, nec Sturm, 1815 "Normandia"
~sculptipennis Sirguey, 1931, nec Chaudoir, 1877 "Normandia"
~viridicollis Sirguey, 1931, nec Kraatz, 1882 "Normandia"
~helvetiae Breuning, 1932; un; nom.pro pauli Csiki, 1927 Switzerland:Emmenthal
=armoricanus Oberthür, 1935
=melanipes Lienhart, 1936
~fuliginosus Frennet, 1937, nec Panzer, 1809
=garactensis Alluaud, 1937
~mormalensis Leleup, 1938; un
~bourgini Balazuc, 1946; un
=duponti Lebis, 1949 =beskydicus Smetana, 1949 N-Moravia:Moravskoslezské Beskydy Mts.(Lysá hora Mt.)
=lueneburgensis Blumenthal, 1949 N-Germany:Niedersachsen:Lüneburger Heide
=pseudoviridipennis Lebis, 1949
~fritzschei Korell, 1950; un
~edmundi (Lebis, 1955); un
~vulcani (Lebis, 1955); un
=syrmiensis Mandl, 1955 Serbia:Frushka Gora Massif(?error)
~christyae Dorsselaer, 1957; un
~schwarzwaldensis Mandl, 1962; un Schwarzwald:Goldersbach, 1000m
~subignifer Mandl, 1967; un SC-Germany:Baden:Holzenschallbach nr.Lörrach
~pumicatoides Mandl, 1968; un
~pseudoceroglossoides Tarrier, 1975; nud
~martinae (Culot, 1978); un
~hennuyi (Grafteaux, 1980); un
~chlorochromus (Culot, 1981); un
~chrysocyaneus (Culot, 1981); un
~deledecquei (Culot, 1981); un
=forsteri Mandl, 1981 Austria:Niederosterreich:St.Aegyd:Mt.Göller(=betw.Sankt Aegid/Mariazell)
~grafteauxi (Hennuy, 1981); un
~auverloti (Thibaudeau, 1983); un
~thumseri (Thibaudeau, 1983); un
~inopinatus (Dachy, 1986); un
~rubicundosus (Branger, 1986); un
~noviprati Maquet, 1991; un
~atronitens Henderickx, 1998; un ?Li?ge env.
~bampsi Henderickx, 1998; un ?Li?ge env.
~spinolatus Maquet, 1999; un Belgium:Li?ge env.

ssp. cupreonitens Chevrolat, 1861 NW-France:Calvados:Ft.de Cerisy, Ft.de Ecouves
~fastuosus Le Moult, 1912; un ?Ft.de Cerisy
~pseudomelas Le Moult, 1912; un ?Ft.de Cerisy
~pseudopurpureus Le Moult, 1912; un ?Ft.de Cerisy
~roeschkei Le Moult, 1912; un ?Ft.de Cerisy
~rossii Le Moult, 1912; un ?Ft.de Cerisy
~cauvini Sirguey, 1931; un Calvados:Ft.d’Ecouves
~tourteani Sirguey, 1931; un Calvados:Ft.d’Ecouves
~pseudonigripes Machard, 1982; un
~nigronitens (Lacroix, 1986); un

ssp. costellatus Géhin, 1882; ?non priority C,W-France:Lot - Lozere - Cantal - Allier - Finistrére - Cotes-du-Nord; "Auvergne"
=minor Kraatz, 1860; ?un; ?priority
=bleusei Oberthür, 1884 "Bretagne:Foret de Lorges"
~melas Oberthür, 1884, nec Creutzer, 1799 "Bretagne:Foret de Lorges"
~purpureus Oberthür, 1884, nec Kraatz, 1876 "Bretagne:Foret de Lorges"
=subfestivus Oberthür, 1884 Finistrére:Foret de Lorges
~viridipennis Bleuse, 1896, nec Lewis, 1880
=semicupreonitens Schulz, 1900
=laevicostatus Schulz, 1900
~guerryi Born, 1907, nec Born, 1903 "Auvergne:Pic de Sancy"
=quittardi Lapouge, 1908 ?Dep.Cantal
=bonneti Le Moult, 1912 Puy-de-Dome:Mt.Doré
~clermonti Le Moult, 1912; un
~cupreicollis Le Moult, 1912; un
~logreensis Le Moult, 1912; un
~mesmini Le Moult, 1912; un
~nigriformis Le Moult, 1912; un Puy-de-Dome:Mt.Doré
~coeruleus Barthe, 1924; un Rennes env.
=buyssoni Barthe, 1929 Allier:Moulins env.
~cupreus Oberthür, 1935, nec Linné, 1758
~leleupi Hansen, 1968; un
~donnoti (Coupat, 1971); un
~lequeti (Darnaud, 1978); un
~artusi (Le Mao, 1984); un
~jacquesi (Lacroix, 1984); un
~quinti (Lacroix, 1984); un
~coatlochensis (Branger, 1986); un
~igneus (Branger, 1986); un
~nigrescens (Branger, 1986); un
~damieni (Lacroix, 1986); un
~delphinae (Lacroix, 1986); un
~nigrinofemoratus (Lacroix, 1986); un
~nigronissimus (Lacroix, 1986); un
~ambazacensis (Dralet, 1999) Hte.Vienne:N of Ambazac

ssp. kraussi Lapouge, 1898; ?non priority NE-Italy; S,E-Austria; W-Hungary:Vas:Köszeg; ?LNF
=intercostatus Gredler, 1854; ?un; ?priority "Passeiertal"(=NE-Italy)
=vindobonensis Kubik, 1901 NE-Austria:Wienerwald Mts.; ?Wien

ssp. escheri Palliardi, 1825 E-Europe:Carpathians:NW-Slovakia to Banat; "Banat"
=opacus Haury, 1878 "Transsylvania"
=rugosipennis Géhin, 1882 "Transsylvania"
=fussi Birthler, 1886
~laevipennis Seidlitz, 1891; un
=istratii Hormuzaki, 1901 "Bukowina"
~laetus Lapouge, 1902; un "Serbia"(error)
~funestus Csiki, 1905; un "Transsylvania"
=decebali Mallasz, 1929 "Siebenburgen"
~interruptecostatus Marcu, 1934; un
~palustris Niedl, 1946; un ?N-Slovakia:Vysoké Tatry Mts.:Tatranská dolina valley
~cyaneipennis Breuning, 1947; un "Russ.subc.:Vihorlat"(=E-Slovakia:Vihorlat Mts.)
~pseudoputzeysi Breuning, 1947; un "Rumänien:Szamosujvar"
~carpaticus Niedl, 1948; un W-Ukraine:Uzhgorod
~contortus Niedl, 1948; un
~diffisus Niedl, 1948; un W-Ukraine:?Uzhgorod
~havelkai Niedl, 1948; un
~kulti Niedl, 1948; un W-Ukraine:Uzhgorod
~latissimus Niedl, 1948; un W-Ukraine:Jasina
~napravniki Niedl, 1948; un W-Ukraine:?Mukachevo
~punctatus Niedl, 1948; un W-Ukraine:?Uzhgorod
~mandli Niedl, 1960; un; nom.pro carpaticus Niedl, 1948 W-Ukraine:Uzhgorod
~nitidis Vicol, 1987; un
~takacsi Vicol, 1987; un
~viridis Vicol, 1987; un

ssp. festivus Dejean, 1826, nec Panzer, 1796; nbm SE-France:Aude:N-part; Tarn:S-part(Soréze)
~holochrysus Barthe, 1908; un
~nigrofemoratus Barthe, 1908; un
~purpureorutilans Barthe, 1908; un
~violaceopurpureus Barthe, 1908; un
~castaneipes Lapouge, 1910; un
~crassepunctus Lapouge, 1910; un
~holochrysum Lapouge, 1910; un
~pumicatus Lapouge, 1910; un
~nigricollis Le Moult, 1912; un
~dauphini Auzat, 1915; un "Foret de la Pauze"
~ceroglossoides Barthe, 1924; un
~auratopumicatus Sirguey, 1924; un
=charpentieri Sirguey, 1931
=gauberti Sirguey, 1931
=lebisi Sirguey, 1931
~claripes (Alabergere, 1968); un
~mirilius (Alabergere, 1968); un
~aveyronnensis (Raynaud, 1970); un Aveyron:?
~jeanimarcianus (Culot, 1981); un
~benedictae (Lacroix, 1986); un
~nigrocyanescens (Devecis, 1988); un
~cyaneonigrescens (Devecis, 1991); un

ssp. punctatoauratus Germar, 1826 SC-France:Pyrenees-Orientales("Mts.E of Mont Louis"-NT); Ariége; "Pyrenees"
=canigouensis Schaefer, 1966 Pyrenees-Orientales:Canigou Massif:Pla Guilhem
=romani (Raynaud, 1968)
=roussellei (Colas, 1969)
=errensis (Rousselle, 1974) Pyrenees-Orientales:Vallée d’Err:NW-slopes betw.Puigmale/Pic du Pas dels Lladres
~holoviridis (Devecis, 1988;) un
~guerroumii Leplat, 1989; un Ft.de Nohédes(?Pyren.Orientales)

ssp. farinesi Dejean, 1826 SC-France:Pyrenees-Orientales, Aude, Ariege
=carlittensis Barthe, 1910 ?Pyrenees-Orientales
=pseudofestivus Lapouge, 1910 Ariege:Ft.de Bosc
=barthei Lapouge, 1912 Aude & Ariege:N-part:Beclaire env.& Ft.de Belesta
=boscensis Nicolas, 1923 Ariege:Ft.de Bosc
=bouisseti Barthe, 1936
=aubryi (Colas, 1969) Ariége:Cirque d’Anglade nr.Salau
~ninini (Colas, 1969); un Ft.de Gravas
=raynaudi (Colas, 1969) ?Pyren.Or.:Ft.de Boucheville
=angladensis (Jeanne, 1969) Ariége:Salau:cirque d’Anglade, 1400m
~vallierensis (Jeanne, 1969), nec Colas, 1964 Ariége:Mt.Vallier Massif:Couflens:Arreou lake, 1800m
~simardi (Raynaud, 1970); un
=pailherensis (Jeanne, 1972) Ariége:Port de Pailhéres, 2000m
=pseudobarthei (Schaefer, 1973) Aude:Massif S of Fajolle
~ayensis (Darnaud, 1980); un Ariége:Ayens
~caroli (Darnaud, 1980); un Ariége:Col des Croisets
~costessequi (Darnaud, 1980); un Ariége:Ft.du Castera
~philippi (Darnaud, 1980); un Ariége:Port de Pailhéres
~tarasconensis (Mercier, 1988); un
~sylvius (Delhaye, 1988); un
~mauricola Leplat, 1998; un E-Pyrenees:Serra de Mauri, Camporells Road

ssp. montanus Géhin, 1882; ?non priority Hautes-Pyrenées(Gavarnie); Haute Garronne; Pyren.Atl.:E-part; NE-Spain:Lerida:C-Pyrenees
=flavolimbatus Géhin, 1876; ?un; ?priority
~lugubris Géhin, 1881, nec Duftschmid, 1812 Hautes-Pyrenées:Pic de Néré
=pandellei Géhin, 1882 Haute-Garonne:Bagneres de Luchon
~rubripes Nicolas, 1898, nec Duftschmid, 1812 Hautes-Pyrenées:Pic de Néré
=aurocostatus Lapouge, 1902 ?Hte-Garonne:Bagneres de Luchon(?inexact: ?in fact:NE-Spain:C-Pyrenees:Val d’Aran env.)
=bigerrio Lapouge, 1924 Hautes-Pyrenées:Baréges env.
~illixioni (Darnaud, 1981); un Haute-Garonne:"Haute montagne de Luchon"



2.lineatus Dejean, 1826 N-Spain, N-Portugal, W-Pyrenees

ssp. lineatus Dejean, 1826 N-Spain:Oviedo:E-part to Navarra:Pamplona; Logrono; C-Spain:Avila; "Asturia"
=basilicus Chevrolat, 1837 "Porto Rico"(error)
=whitei Deyrolle, 1852 LNF(unlabelled HT)
~debilicostis Kraatz, 1896, nec Kraatz, 1878 "Asturia"
~forticostis Kraatz, 1896, nec Kraatz, 1878 "Asturia"
~obscurus Le Moult, 1913; un Asturia:Puerto Pajares
~rufipes Lapouge, 1931; un
~fuliginosus (Puisségur, 1962), nec Panzer, 1809 ?Sierra de Andia
~avilensis Breuning, 1966; un Picos de Europa Massif:Aliva refuge(=Avila error)
~holoidus (Raynaud, 1969); un Sierra de Andia
=numancianus (Raynaud, 1969) Soria:Duero River source area
~poussini Bra?es, 1988; un Palencia:Revila env.(Cervera de Pisuerga)

ssp. troberti Kraatz, 1860 NC-Spain:Guipuzcoa; Navarra:NW-part; SW-France:Pyrenees-Atlantiques
=ursuius Lapouge, 1911 SW-France:Pyrenees-Atlantiques:Mt.Ursuia
=beharrius Nicolas, 1916 Pyr.Atlantiques:Mt.Beharria S of St.Jean Pied de Port
=hochstetteri Born, 1919 Pyr.Atlantiques:Val d’Urepel
~incostatus Nicolas, 1919; un
=miniatulus Venet, 1930 Pyr.Atlantiques:Val d’Urepel

ssp. lateralis Chevrolat, 1840, nec Fabricius, 1792; nbm N-Portugal; NW-Spain:La Coruna, Orense, Pontevedra, Lugo, Leon, Oviedo; "Sra.de Tranquera"
~bilineatus Géhin, 1885; un "Portugal"
=strasseri Lauffer, 1905 Leon:Laceana
=martinezi Bolivar, 1922 Leon:Foncebados
=leonensis Born, 1918 Leon:Ponferrada
~subholoviolaceus (Puisségur, 1962); un Galicia:Sierra de Arbolente
~glaucus (Raynaud, 1969); un LNF
~holochlorus (Raynaud, 1969); un LNF
~quasilineatus Tarrier, 1975; un Oviedo:Alto del Connio
~annikae (Darnaud, 1979); un Leon:Ponferrada
~boulbenei (Darnaud, 1979); un Leon:Ponferrada
~bourdeaui (Darnaud, 1979); un Leon:Ponferrada
~caubeti (Darnaud, 1979); un Leon:Ponferrada
~daniellenae (Darnaud, 1979); un Leon:Ponferrada
~rothei Bra?es, 1985; un
~sanabriensis Bra?es, 1985; un Zamora:Sanabria Lake
~littoralis Bra?es, 1988; un Pontevedra
=petrimeyeri Maux, 2002 Leon:nr.Puerto de Monteviejo, 1400m

ssp. salmantinus Bolivar, 1922 CW-Spain:Salamanca/Caceres:Sierra de Gata; Salamanca:Sequeros
~subsalmantinus Bra?es, 1985; unCaceres:St.Martin de Trevejo
=sainvali Bra?es, 1987 Caceres:Hervas



3.olympiae Sella, 1855 N-Italy:Piemonte:Biella env.:Sessera valley
~sellae Stierlin, 1881, nec Kraatz, 1878
=stierlini Heyden, 1881; nom.pro sellae Stierlin
~fucatus Lapouge, 1902; un



4.solieri Dejean, 1826 Alps of SE-France & NW-Italy

ssp. solieri Dejean, 1826 SE-France:Alpes-de-Haute-Provence(?"Basses-Alpes:Les Dourbes"), Alpes-Maritimes
~furcillatus Bleuse, 1913; un "Basses-Alpes"(=Alpes-de-Haute-Provence)
~louveti Clermont, 1922; un Alpes-Maritimes:St.Martin de Vesubie
=fuscoaeneus Clermont, 1925
=vesubiensis (Poncetton & Reymond, 1935) Alpes-Maritimes:St.Martin de Vesubie
~fagniezi Colas, 1959; un Alpes-Maritimes:St.Martin de Vesubie
=cavalieri (Colas, 1965) Alpes-Maritimes:Col de Braus(=nr.Sospel)
=altamontanus (Ochs, 1965) Alpes-de-Haute-Provence/Alpes-Maritimes:Col de la Cayolle
=chalcostola (Ochs, 1965) Alpes-Maritimes:Collongues
=tarrieri (Ochs, 1965) Alpes-Maritimes:Lantosque
~ater (Raynaud, 1970); un
~gordolasquensis (Raynaud, 1970); un Alpes-de-Haute-Provence:Gordolasque valley
~smaragdinus (Raynaud, 1970); un Alpes-Maritimes:St.Martin de Vesubie
~viridicupreus (Raynaud, 1970); un
~florettae Raffaldi, 1971; un Alpes-de-Haute-Provence:Annot env.
~hyperchlorus Tarrier & Bellon, 1972; un Alpes-Maritimes:St.Martin de Vesubie
~piezus Bellon & Tarrier, 1972; un Alpes-de-Haute-Provence/Alpes-Maritimes:Col des Champs
~monicae Tarrier, 1973; un Alpes-Maritimes:Lantosque
~carreti Tarrier, 1975; un Alpes-Maritimes:St.Etienne-de-Tinée
~desmon Bellon & Tarrier, 1975; un Alpes-Maritimes:Belvédere
~dujardini Bellon & Tarrier, 1972; un Alpes-Maritimes:Sospel
~rouyricus Tarrier, 1975; un Alpes-de-Haute-Provence:Mt.Vial

ssp. clairi Géhin, 1885 NE-Italy & SE-France:Alpes-Maritimes(?"Mercantour"; ?Col de Tende; ?error)
=cyaneoviridis Carret, 1897
~azurescens (Raynaud, 1970); un
~viridiater (Raynaud, 1970); un
~axiomorphus Tarrier, 1971; un Alpes-Maritimes:La Muselle
~devillei Tarrier, 1972; un
~strobinoi Tarrier, 1975; un Alpes-Maritimes:"Bas vallon de Molliéres:Pierre Blanche"

ssp. liguranus Breuning, 1933 NW-Italy:Alpi Liguri(Mte.San Giorgio) & SE-France:Alpes-Maritimes:Tende env.
~sturanii (Raynaud, 1970); un LNF

ssp. bonnetianus Colas, 1936; nom.pro bonneti Colas SE-France:Var:E,NC-part(St.Cassien des Bois); Alpes-Marit.; Alp.Hte.Prov.:Teillon Massif
~bonneti Colas, 1936, nec Le Moult, 1912 Var:St.Cassien des Bois
=bonadonai Colas, 1947 Alpes-Maritimes:Séranon
=canjuersicus (Ochs, 1965) Var:NC-part:Ft.Grand Margés:Aiguines
=curtii (Ochs, 1965) Alpes-Maritimes:Vescous
=belloni (Tarrier, 1966) Alpes-Maritimes:Roquesteron
=caussolsicus (Bellon, 1971) Alpes-Maritimes:Caussols env.
~teillonensis Tarrier, 1975; un Alpes-de-Haute-Provence:Mt.de Teillon



5.splendens Olivier, 1790 S-France:Aveyron-Pyr.Atl.-Landes; NE-Spain:Lerida, Huesca, Navarra, Alava; Jamaika(error)
=carolinus Fabricius, 1792 Carolina(error)
=viridis Dejean, 1826
=cychricollis Lapouge, 1898 Haute-Garonne:Luchon
~lineatopunctatus Lapouge, 1898, nec Dejean, 1826
~ovipennis Lapouge, 1902; un
~seriepunctatus Lapouge, 1902; un
=nereensis Nicolas, 1902 Hautes-Pyrenées:Pic de Néré
~vittatus Lapouge, 1902; un Aude:Ft.de Fanye
~mordoratus Nicolas, 1908; un Hautes-Pyrenées & Haute-Garonne:Lourdes & Luchon
~jeani Clermont, 1909; un Ft.de Lestelas
=ammonius Lapouge, 1909 Tarn:Montagnes Noires
=pinetorum Lapouge, 1910 "Dep.Landes"
=pseudotroberti Lapouge, 1910
~babaulti Le Moult, 1912; un Tarn:Montagnes Noires
~goliath Le Moult, 1912, nec Morawitz, 1886 Aude:Ft.des Fanges
~granulipennis Le Moult, 1912; un Tarn:Montagnes Noires
=pseudocychricollis Le Moult, 1912 Tarn:Montagnes Noires
=pseudoovipennis Le Moult, 1912 Tarn:Montagnes Noires
=pseudoviridicollis Le Moult, 1912 Aude:Ft.des Fanges
=pseudovittatus Le Moult, 1912 Aude:Ft.des Fanges
=subcychricollis Le Moult, 1912 Aude:Ft.des Fanges
=subovipennis Le Moult, 1912 Aude:Ft.des Fanges
~vinosus Le Moult, 1912; un Tarn:Montagnes Noires
~viridicollis Le Moult, 1912, nec Kraatz, 1882
~dombrowskii Le Moult, 1913; un Haute-Garonne:Luchon
~laticollis Le Moult, 1913; un Pyrenees:Lerida:Val d’Aran
=piesbergeni Le Moult, 1913
=lapurdanus Lapouge, 1913?"Basses Pyrénées:S-part of Bayonne district"(=Pyr.-Atlantiques); ?Pyr.-Atl.:Cambo
=mascarauxi Lapouge, 1916 "Basses Pyrénées:Vallée de la Bidassoa & Vallée se la Nivelle"(=Pyr.-Atlantiques)
~cervicis Csiki, 1917; un; nom.pro laticollis Le Moult Pyrenees:Lerida:Val d’Aran
~viridicervix Csiki, 1917; un; nom.pro viridicollis Le Moult
~pallens Lapouge, 1924, nec Fabricius, 1775 "Basses Pyrénées"(=Pyr.-Atlantiques; alpine form)
=hemipterus Sirguey, 1931
=palesius Sirguey, 1931 "Pic Burat"
~cardinalis (Raynaud, 1968); un
~najacensis (Raynaud, 1968); un Aveyron:Ravin de Najac
~temnoglypha (Raynaud, 1969); un
~ursuiensis (Raynaud, 1969); un ?Pyrénées-Atlantiques:Mt.Ursuia
~violaceocyaneus (Jeanne, 1972); un Aude:Ft.d’Arques
~arradoyensis (Lequet, 1975); un Pyr.-Atlantiques:Pic d’Arradoy, 600m
~violaceoviridis (Rouselle, 1978); un
~alberginius (Remond, 1986); un
~michaellae (Remond, 1986); un
~incertus (Devecis, 1988); un
~taffini (Delaporte, 1988); un


BACK



PROCERUS Dejean, 1828; type species Carabus scabrosus: Fabricius, non Olivier = Carabus gigas Creutzer
species: 3



1.gigas Creutzer, 1799 Yugoslavia, Greece, S-Romania, SW,C,NE-Bulgaria, SE-Austria, NE-Italy

ssp. gigas Creutzer, 1799 NE-Italy; SE-Austria, Yugoslavia("Idria, Carniolia"), S-Romania; SW,C,NE-Bulgaria
~scabrosus: Fabricius, 1801, non Olivier, 1795
~limbatus Géhin, 1885; un
~olympicus Lapouge, 1902, nec Géhin, 1885 ?Olympos Massif
~limbiger Csiki, 1927; un; nom.pro limbatus Géhin

ssp. parnassicus Kraatz, 1884 Greece, Macedonia, S-Albania

ssp. duponcheli Dejean, 1831 S-Greece:Peloponnes Peninsula
=kusdasi Mandl, 1966 Peloponnes:Chelmos(alpine form)



2.scabrosus Olivier, 1795 SE-Bulgaria, Krym Penins., Turkey, Caucasus, Georgia, Armenia, Azerbaydzhan, ?N-Iran

ssp. scabrosus Olivier, 1795 SE-Bulgaria, Eur.Turkey("Konstantinopel"=Istanbul)
=olivieri Dejean, 1826 ?"Konstantinopel"
=?aegyptiacus Motschulsky, 1844 "Egypt"(error)
~cribratus Motschulsky, 1850, nec Quensel, 1806 "Constantinople"(=Istanbul; LT)
~angusticollis Haller, 1885, nec Fabricius, 1801
~dejeani Géhin, 1885; un

ssp. propinquus Csiki, 1927; nom.pro giganteus Bodemeyer NW-Turkey:Istanbul env.(Alem Dag SE of Istanbul), Adapazari
~giganteus Bodemeyer, 1915, ?nec Heer, 1837 Alem Dag SE of Istanbul

ssp. mentor Blumenthal & Breuning, 1967 NW-Turkey:Izmit, Adapazari, Bolu(Bolu Mts. N of Mengen)

ssp. schuberti Breuning, 1968 NC-Turkey:SW of Sinop:Cangal Dag; Zonguldak, Cankiri, Kastamonu, Sinop

ssp. culminicola (Cavazzuti, 1989) NW-Turkey:Bursa:Ulu Dag:alpine zone, 2000-2150m

ssp. tauricus Bonelli, 1811 Ukraine:Krym Peninsula; "Crimea meridionale"
=aeneus Motschulsky, 1850 "Tauria"(LT)
=nigritulus Kraatz, 1876
=purpureus Kraatz, 1876
=viridissimus Kraatz, 1876

ssp. caucasicus Adams, 1817 Caucasus:N-part("Konstantinogorsk")
~tuberculosus Géhin, 1885; un

ssp. audouini Brullé, 1835 NE-Turkey:Samsun to Rize(Trabzon env.)
~obscurus Géhin, 1885; un

ssp. colchicus Motschulsky, 1844 Caucasus:S-part(Oni); Georgia; NE-Turkey:Artvin
=shavsheticus Zaitzev, 1930 NE-Turkey:lower Çoruh River

ssp. armenius Zaitzev, 1930 NE-Turkey:Bayburt:Gümüshane env.(betw.Gümüshane/Erzincan)

ssp. ispiratus Cavazzuti, 1986 NE-Turkey:Erzurum:Ispir env.(upper Çoruh Nehri valley 5km W Ispir)

ssp. fallettianus (Cavazzuti, 1997) SE-Armenia:Zangezur Mt.R.:Khndzoresk, Megri env.

ssp. myskai Cavazzuti, 2004 SE-Armenia:Kafan env.

ssp. boulbenarlettae Cavazzuti & Myška, 2004 NE-Azerbaydzhan:Quba, 800m; 1m

ssp. farnazae Deuve, 2006 NW-Iran:Arasbaran:Kalibar:Makidi, 1650m(~38:50N/46:55E); 1m/1f

ssp. sommeri Mannerheim, 1844 NW-Turkey:Bursa, Bilecik; "Brussa"(=Bursa)
~bosphoranus Motschulsky, 1844, nec Fischer, 1823 ?Bursa
=breviusculus Kraatz, 1876 Bursa
~uludagensis Schweiger, 1964; un Uludag nr.Bursa:Incaya village env.:chestnut forest
=basoglui Schweiger, 1964 Uludag nr.Bursa:fir forest, 1700m

ssp. dardanellicus Kraatz, 1886 NW-Turkey:Çanakkale; Asiatic coast of Dardanelles Strait

ssp. sterilis Bodemeyer, 1915 NW-Turkey:Bilecik:Boz Dag

ssp. amasicus Csiki, 1927; nom.pro modestus Kraatz NC & C-Turkey(Amasya)
~modestus Kraatz, 1876, nec Fabricius, 1801 Amasya

ssp. bureschianus Breuning, 1928 SC-Bulgaria:Rodopes Mt.R.(Backovo), NE-Greece

ssp. ~montisabanti Schweiger, 1962; un NW-Turkey:Bolu:S of Abant(alpine steppes S of Abant)

ssp. callipygius Cavazzuti, 1986 NC-Turkey:Kastamonu(karstic plains ca.20km N Devrekani); Cankiri
=cerkes Basquin & Darge, 1986 Cankiri:Ismetpaºa env.

ssp. munzurensis Cavazzuti & Lassalle, 1987 CE-Turkey:Tunceli:Pülümür env.(pass N of Pülümür)

ssp. estegeicus (Cavazzuti, 1989) SW-Turkey:(Isparta:Davraz Dag)

ssp. weidneri (Lassalle, 1990) SC-Turkey:Kahramanmaras:Göksun, Cardak; Adana:Saymbeyli

ssp. bulgharmaadensis Bodemeyer, 1915 S-Turkey:Konya:Bolkar Dag:N-slope("Bulghar-Maaden"), Eregli:Ivriz env.; Mersin:Namrun

ssp. transversalis Csiki, 1927, nec Duft.; nbm; nom.pro laticollis Kraatz SC-Turkey:Mersin("Külek"=Gülek), Adana
~laticollis Kraatz, 1876, nec Sturm, 1826 Mersin:"Külek"(=Gülek)
=toulgetianus Deuve, 2002; nom. pro transversalis Csiki Mersin:"Külek"(=Gülek)

ssp. akbesianus Breuning, 1975; non junior homonym SC-Turkey:Antakya(Akbes nr.Iskenderun), Adana, Mersin, Kahramanmaras
=akbesiensis Toulgoet, 1987; nom.pro akbesianus Breuning Antakya:Akbes nr.Iskenderun

ssp. kahramanicus Cavazzuti & Myška, 2004 SC-Turkey:Kahramanmaraº:Kahramanmaraº env.

ssp. elbursianus Mandl, 1958 N-Iran:Elborz Mt.R.:Taht-i-Suleiman Massif:Sardab/Hasankif valley(~36:30N/51:10E); 1f



3.syriacus Kollar, 1843 SE-Turkey, W-Syria, N-Israel, Lebanon

ssp. syriacus Kollar, 1843 SE-Turkey:Antakya env.("Djebel Okra"=Ziyaret Dagi); NW-Syria
=daphnensis (Korell, 1985) Antakya:mts.24km S Harbiye

ssp. galilaeus Schweiger, 1970 N-Israel(Galilea:Elon); S-Lebanon; SW,CW-Syria

ssp. limitaneus (Korell, 1989) SE-Turkey:Antakya:Reyhanli env.(2km S Reyhanli); NW-Syria:Daret’Azze

ssp. phoeniceus Deuve, 2004 N-Lebanon:Kesrouane:Wata-al-Jaouz


BACK



MEGODONTUS Solier, 1848; type species Carabus caelatus Fabricius
species: 20



1.avinovi Semenov & Znojko, 1932 Sakhalin(Sakhalin, central part) & Moneron Island



2.bonvouloiri Chaudoir, 1863 NE-Turkey

ssp. bonvouloiri Chaudoir, 1863 NE-Turkey:Giresun, Trabzon env., Rize; Gümüshane:SE of Zigana Pass
~clandestinus Schaufuss, 1882; un
~foveicollis Schaufuss, 1882; un
~nobilis Schaufuss, 1882; un
~pastillatus Lapouge, 1902; un
~kaschkarensis Blumenthal & Breuning, 1967; un Rize:Kaçkar Dagi(Ardesen valley:Ayder); Erzurum:Ispir, Çoruh valley, Ovit Pass
~kolatdaghensis Breuning, 1967; un "Kolat Dag nr.Trabzon, 2000m"(=Mts.nr.Zigana Pass)
=soganlidaghensis Machard, 1991 Trabzon:Soganli Pass S of Çaykara, 2300m

ssp. graciliformis Breuning, 1964 NE-Turkey:Ordu:Akkus(forest zone nr.Akkus, 1300-1500m), Gölköy env.

ssp. fortunatus Heinz, 1970 NE-Turkey:Artvin(pass N of Murgul, 1000m) & Rize:deciduous forests along Black Sea coast

ssp. casalei Cavazzuti, 1986 NE-Turkey:Artvin:Göktas valley



3.caelatus Fabricius, 1801 Slovenia, Bosnia, Croatia; N,C-Albania; Italy:Trieste env.

ssp. caelatus Fabricius, 1801 C,E-Slovenia; ?E-Austria:Karawanken; Bosnia; E-Montenegro; N,C-Albania; "Krain"
=carniolicus Crotch, 1871
=sarajevoensis Apfelbeck, 1890 Bosnia:Sarajevo env.
=volujakianus Apfelbeck, 1894 Bosnia:Volujak Massif; alpine form
=hilfi Born, 1907 Bosnia:Vran Planina & Cvrstnica Planina; alpine form
=malissorum Apfelbeck, 1919 ?Montenegro:Durmitor; alpine form
=metalkanus Apfelbeck, 1919 SE-Bosnia:"Metalka Pass"(=SE of Gorazde)

ssp. dalmatinus Duftschmid, 1812 Adriatic Sea Coast:Rijeka to N-Albania; ?LNF
=macretus Kraatz, 1877 Dinara Planina; alpine form
~ljubinjensis Haury, 1885; un Bosnia:Ljubinje
=procerus Reitter, 1885 "W-Montenegro"
=cabuljensis Apfelbeck, 1919 Bosnia:Mostar env.:Cabulja Planina; alpine form
=dinaricola Müller, 1930 ?"Dinarische Alpen"(=Dinara Planina)

ssp. schreiberi Kraatz, 1887 Italy:Trieste; SW-Slovenia("St.Canziano, Schneeberg=?Sneznik), W-Croatia; NW-Bosnia
=grmecensis Born, 1910 Bosnia:Grmec Planina



4.croaticus Dejean, 1826 Slovenia, Croatia, Bosnia, Montenegro, N-Macedonia, N-Albania

ssp. croaticus Dejean, 1826 S-Slovenia, W-Croatia & W-Bosnia:Adriatic coast mts.; Croatia:"Ostaria btw.Gospic/Carlopago"
~carniolicus Géhin, 1885; un ?"Krain:Mt.Nevoso"
=schmidti Apfelbeck, 1890 Bosnia:"Vucja Luka nr.Sarajevo"
=primarius Lapouge, 1902 Croatia:Velebit Mts.
=frankenbergeri Obenberger, 1914 NW-Croatia:"Gr.Capella"(=Velika Kapela Mts.)
~mediterraneus Apfelbeck, 1919, nec Born, 1906 Bosnia:Cabulja Planina nr.Mostar
~carnicus Csiki, 1927, nec Depoli, 1925; nom.pro carniolicus Géhin ?"Krain:Mt.Nevoso"
=fodori Csiki, 1927; nom.pro mediterraneus Apfelbeck Bosnia:Cabulja Planina nr.Mostar
=albiensis Depoli, 1938; ?nom.pro carnicus Csiki ?"Krain:Mt.Nevoso"

ssp. bosnicus Apfelbeck, 1890 C,N-Bosnia(Treskavica Mts.)
=kobingeri Apfelbeck, 1904 C-Bosnia:"Mts.near Foinica & Travnik"
~travnikanus Apfelbeck, 1904; un C-Bosnia:?Travnik
~leonhardianus Breuning, 1932; un C-Bosnia:Cvrstnica Planina
=pretneri Krätschmer & Drovenik, 1977 N-Bosnia:Ostrelj env.(Ostrelj, 1000m), Srnetica Mts.

ssp. zepcensis Reitter, 1902 E-Bosnia:Zepce env.; Ravan, Zvijezda, Javor, Ozren Mts.; CW-Serbia:Murtenica planina
=bosiljevici Drovenik & Paviæeviæ, 1985 CW-Serbia:50km S Titovo Uzice:Murtenica planina:Brijac

ssp. durmitorensis Apfelbeck, 1904 Montenegro(Durmitor Massif) & NE-Albania:Prokletije Mts.; Kosovo; N-Albania, W-Maced.
=babinjensis Apfelbeck, 1906 Montenegro & NE-Albania:Prokletije Mts.:"Babinje Planina"
=ljubetensis Apfelbeck, 1918 Serbia/Macedonia/Albania borders region:Sar planina Mts.:Ljubeten
=droveniki Krätschmer, 1984 SW-Macedonia:Galicica Mts., Baba Mts., Pelister
=kraetschmeri Drovenik, 1978 SE-Montenegro:betw.Ivangrad/Rozaj:Turjak



5.danilevskii Obydov, 1993 W.Sayans:S of Abakan(left Yenisey bank:Dzhoysky P.nr.Cheryomushki); ?W-Tuva:Barlyk R.
=viridicans Obydov, 1993 West Sayans:S of Abakan:Borus Mt.R.:Borus Pass, 800-1000m



6.ermaki Lutshnik, 1924; nom.pro amoenus Chaudoir; ?non priority arctic Eur.Russia; N-Siberia; Altays, Sayans
~amoenus Chaudoir, 1852, nec Baudet-Lafarge, 1836 Altays
=kantaikensis Géhin, 1885; ?un; ?priority ?"Kantaika"(=arctic W-Siberia:SW of Noril’sk)
=laetensis Obydov, 2000 NW-Siberia:Komi:Usa:Lek-Rogovaia River mouth



7.exaratus Quensel, 1806 N-Caucasus:Anapa to Mt.Kazbek; Krasnodar:Taman Pen.to Manych R..; "Kislar"(=?Kizlyar)
~?seriatus Motschulsky, 1850, nec Wiedemann, 1821; ?hyb ?Balkaria; ?hyb exaratus/violaceus aurolimbatus
=multicostis Reitter, 1888 ?Circassia
=septemlineatus Reitter, 1888
=prahwei Lutshnik, 1909 ?Dagestan
=lebedewi Lutshnik, 1927; nom.pro seriatus Motschulsky; ?hyb ?Balkaria; ?hyb exaratus/violaceus aurolimbatus
=ordinatus Csiki, 1927; nom.pro seriatus Motschulsky; ?hyb ?Balkaria; ?hyb exaratus/violaceus aurolimbatus
=exaratulus Beheim & Breuning, 1943 W-Caucasus:Mt.Fisht



8.gyllenhali Fischer, 1827, nec Paykull, 1790; nbm; nom.pro dejeani Fischer Krym Peninsula
~dejeani Fischer, 1823, nec Fischer, 1822 Krym Peninsula
=blakistoni Newman, 1857; ?priority



9.imperialis Fischer, 1823 Altays:Semipalatinsk, Ust’-Kamenogorsk, upper Irtysh River; Barnaul(?error)



10.kolbei Roeschke, 1887 Hokkaido; N,C-Kurils

ssp. kolbei Roeschke, 1887 C-Hokkaido:Hidaka:Niikapu env.; "China"(error)

ssp. aino Rost, 1908 C,E-Hokkaido:Taisetsu Mts., Nemuro, Kushiro; Kurils:Kunashir Island; Hokkaido
=hidakamontanus (Ishikawa, 1966) Hidaka Mt.R.:Tottabetsugawa River Defile
=nitidipunctatus (Ishikawa, 1966) Kamikawa:Shimukappu
=yubariensis (Ishikawa, 1966) Hidaka/Sorachi:Yubari Mt.R.:Mt.Yubaridake

ssp. kosugei (Nakane, 1955) SW-Hokkaido:Otaru, Sapporo env.; ?LNF

ssp. chishimanus (Nakane, 1961) S,C-Kurils:Iturup, Ketoy Island; "Etorofu"(=Iturup)

ssp. futabae (Ishikawa, 1966) NW-Hokkaido:Soya(Nakatonbetsu), Rumoi

ssp. hidakamontanus (Ishikawa, 1966) C-Hokkaido:Hidaka & Tokachi:Hidaka Mt.R.(Tottabetsugawa River)

ssp. kuniakii (Ishikawa, 1971) SW-Hokkaido:Shiribeshi:Niseko Mts.(Mt.Iwaonupuri)

ssp. hanatanii Imura, 1991 Rishiri Island(Mt.Rishiri-daké)

ssp. taikinus (Ishikawa, 1993) SC-Hokkaido:Tokachi:Taiki env.(Taiki-cho:Oshibanosawa)

ssp. mitsumasai Imura, 1994 CW-Hokkaido:Ishikari & Runoi:SW-Mashiké Mt.R.:Hattari, Bokusanai, Yasusé, Bishabetsu

ssp. ushishirensis Obydov & Saldaitis, 1996 C-Kurils:ca.30km NE Ketoy Island:Ushishir Island:Yankicha:Kraternaya:N,SE-part

ssp. munakataorum (Ishikawa, 1969) SW-Hokkaido:Oshima/Hiyama:Daisengen Massif(Mt.Daisengendake)

ssp. yasudai (Ishikawa, 1971) SW-Hokkaido:Hiyama/Shiribeshi:Mt.Karibayama



11.leachi Fischer, 1823 S-Altays

ssp. leachi Fischer, 1823 S-Altays:Zmeinogorsk, Semipalatinsk env.; ?Irtysh River(?HT lost); "Sibiria"(LT)

ssp. panzeri Dejean, 1829, nec Panzer, 1805; nbm S-Altays:Zyryanovsk, Ust’-Kamenogorsk env.; "Sibiria"(HT lost); LNF(unlabelled LT)
=klimenkoi Obydov, 1999 NE-Kazakhstan:E of Ust’-Kamenogorsk:Kalbinsky Mt.R.:Ognevka vill.

ssp. surakovi Obydov, 1995 C-Altays:Shebalino

ssp. sarymsaktensis Obydov, 1999 NE-Kazakhstan:S of Katon-Karagay:Sarym-Sakty Mt.R.



12.obovatus Fischer, 1827 SC-Siberia:Novokuznetsk(Mts.nr.Kuzneck), Kemerovo & E-Novosibirsk Regions; C-Altays
~obovalis Gebler, 1830; iss obovatus Gebler
=pseudoleachi Obydov, 1999 C-Altays:Teletskoe Lake:N-bank(Yailu vill., Artybash vill.)



13.persianus Roeschke, 1896 N-Iran:E-Gilan, W-Elborz Mt.R.

ssp. persianus Roeschke, 1896 N-Iran:Elborz Mt.R.:lower Rudbar valley:Dare Dasht env.; "Elburs"
=damashensis Deuve, 2006 N-Iran:Gilan:Damash, N of Kharehpu, E of Rudbar

ssp. javahardehensis Deuve, 2000 N-Iran:Gilan/Mazandaran:20km WSW Ramsar:Javahar Deh, 36:52N/50:28E

ssp. neydashtensis Deuve, 2000 N-Iran:Gilan:S of Ramsar:Neydasht, 800m

ssp. espeyliensis Deuve, 2001 N-Iran:Gilan:W-Elborz Mt.R.(betw.Espeyli/Siyakhal:Balarud Mts., 450m)



14.planicollis Küster, 1846 C-Romania:S-Carpathians

ssp. planicollis Küster, 1846 C-Romania:Carpathians W of Olt River:Cindrel-Retezat-Godeanu Mts.; ?"Siebenburgen"

ssp. verae Csiki, 1905 C-Romania:Carpathians E of Olt River:Fagaraº to Bucegi Mts.; "Bucsecs"(=Bucegi Mts.)



15.schoenherri Fischer, 1822 CE-European Russia, C-Siberia, Altays, Sayans, N-Mongolia

ssp. schoenherri Fischer, 1822CE-European Russia; SC-Siberia(Barnaul); Altays, Sayans, N-Mongolia
=melanchlorus Fischer, 1827
~gouberti Géhin, 1885; un
=legorskyi Mandl, 1968 East Sayans:Mana River
=sangilenus Obydov, 1997 SE-Tuva:Sangilen Mts.:Balyktyk-Khem River valley:50:17N/96:39E, 2100m

ssp. sajanus Breuning, 1927 West Sayans(Kazyr-Suk River)

ssp. chakassicus Obydov, 1995 Siberia:Hakassia:betw.Krasnoyarsk/Abakan:Saragash env.



16.septemcarinatus Motschulsky, 1840; nom. carinatus MotschulskyS-Caucasus:Mt.Fisht to Lagodekhi; "Kachetia"; Tbatani (LT); Georgia; W-Armenia; NE-Turkey
~carinatus Motschulsky, 1839, nec Duftschmid, 1812 "Kachetia"; Tbatani(LT)
=typhonius Gistl, 1857
=georgicus Géhin, 1885
=fischtensis Reitter, 1889 W-Caucasus:Mt.Fisht
=subexaratus Reitter, 1889
=trigintiduosulcatus Lapouge, 1898
~altitudinorum Breuning, 1934; un; ios; ?hyb
=altitudinum Breuning, 1934; emend.pro altitudinorum Breuning; ?hyb W-Caucasus:Mt.Abago; ?hyb exaratus/septemcarianatus



17.stroganowi Zoubkoff, 1837 Caspian Sea coast: SE-Azerbaydzhan to E-Elborz Mt.R.

ssp. stroganowi Zoubkoff, 1837 NE-Iran:E-Elborz Mt.R.:Gorgan env.("Astrabad"=Gorgan); ?W-Turkmenia:Kopetdag Mt.R.
~galugahensis Lassalle, 2001; un N-Iran:Mazandaran:betw.Behshahr/Gorgan:Mts. S of Galugah, 1800m

ssp. transfuga Semenov, 1898 SE-Azerbaydzhan:Lenkoran,"Talysh"; ?N-Iran:Gilan
=transcurrens Csiki, 1927; nom.pro transfuga Semenov "Talysh"

ssp. morgani Lapouge, 1907 N-Iran:C-Elborz Mt.R.:Khanian nr.Tunekabun(=36:35N/50:49E), Kelardasht, Chalus
=heinrichi Mandl, 1966 C-Elborz Mt.R.:"Pish-kuh"(=?Pishku, 36:41N/50:40E)

ssp. ebrahimii Deuve, 2000 N-Iran:Mazandaran:S of Nur:Ruyan:Kodirsar, 1200m(=?Kudir:36:27N/51:47E)

ssp. mofidii Deuve, 2000 N-Iran:Mazandaran:Kiyasar env.:Vesmine, 1600m(Kiyasar:36:14N/53:22E)

ssp. lavidzhensis Deuve, 2001 N-Iran:Mazandaran:25km SSE Nur:Lavidzh(36:24N/52:00E)

ssp. tiedemanni Breuning, 1934 N-Iran:Gilan Mt.R.("Salansar Dagh"); Gilan:Asalam, Masuleh

ssp. pseudopersianus Deuve, 2001 N-Iran:Gilan:"Dörwögh Dagh", "Tschakit Dagh", "Kiklik Dagh"(=?Rudbar env.)

ssp. luisae Rapuzzi, 2005 N-Iran:Gilan:S of Sayakhal, 36:57/49:51



18.tuvensis Shilenkov, 1996 West Sayans:W-Tuva:80km N Teeli village, Ak-Sug River



19.vietinghoffi Adams, 1812 NW, NC, E-Siberia, Far East, Sakhalin, N-Korea, NE-China; Alaska; NW, ?NC-Canada

ssp. vietinghoffi Adams, 1812 Russia:NW,C,E-Siberia(Lena River N of Yakutsk)
=stschegolewi Poppius, 1906 Lena River
=alaskanus Obydov, 1996 Alaska(Umiat); NW, ?NC-Canada
=obydovi E.Berlov & O.Berlov, 1996 NW-Siberia:Tyumen:Salekhard env.; 1m

ssp. fulgidus Fischer, 1827 Far East; N-Mongolia; "Dauria"
=deminutus Breuning, 1943 NE-China:N-Chingan Mts.:Bachou Lou
=leptoglyptus Obydov, 1995 C-Sakhalin:Tymovsk, Tym River
=bureianus Shilenkov, 1996 Far East:Khabarovsk district:Bureinsky Mt.R.(Suluk River)
=tardokiyanensis E.Berlov & O.Berlov, 1996 Far East:Khabarovsk district:Sikhote-Alin Mt.R.:N-part:Tardoki-Yani Massif; 1f =nazmovi Obydov, 1999 Far East:Khabarovsk:Komsomolskyi Nat.Reserve:Kamennaya Pad’

ssp. bowringi Chaudoir, 1863 Far East:Vladivostok; NE-China:Jilin, Heilongjiang; N-Korea; "Chusan"(=?N-Korea:Chosan)
~schaumi Morawitz, 1862, nec Gaubil, 1849 Vladivostok
=impunctatus Kraatz, 1886 Suyfun River Mouth(=W of Vladivostok)
~nobilis Ganglbauer, 1886, nec Duftschmid, 1812; nom.pro schaumi Mor. Vladivostok (HT lost); "Port Bruce"(=Mys Bryusa, 42:52N/131:28E; LT)
=caesareus Semenov, 1906; nom.pro nobilis Ganglbauer Vladivostok (HT lost); "Port Bruce"(=Mys Bryusa, 42:52N/131:28E; LT)
=moltrechti Semenov, 1907 "Sidemi"(=nr.Vladivostok)
=vietinghoffianus Semenov, 1907 ?Nikolskaya(=Ussuri basin)
=inhumeralis Breuning, 1964 Korea:Mt.Paik-chek-Shan
~borealis (Kwon & Lee, 1984), nec Paykull, 1790 NE-Korea:W of Chongjin:"Mt.Kwanmobong"(=Kwanchang-san)
=lazoensis Obydov, 1999 Far East:Pogranichnyi:Lazovskiy Nat.Reserve:Kordon Amerika
=levadensis Obydov, 1999 Far East:Pogranichnyi:Barabash-Levada village env.



20.violaceus Linné, 1758 Europe:Pyrenees to Scandinavia, Balcans, Caucasus; SW,SC-Siberia, Kazakhstan

ssp. violaceus Linné, 1758 E,S,N-Germany, Scandinavia, Kola Penins., Bohemia, Moravia, W-Slovakia; Brit.Isles; ?LNF
=candisatus Duftschmid, 1812 "Österreich & Böhmen"(=Austria & Bohemia)
=sublaevis Drapiez, 1820 "Russia"(?error)
~bicolor Letzner, 1850; un
~coerulescens Letzner, 1850; un
~cyanescens Letzner, 1850; un
~marginatus Letzner, 1850; un
~nivalis Letzner, 1850; un NE-Bohemia:"Riesengebirge"(=Krkonoše Mts.)
~purpureomarginatus Letzner, 1850; un
~violaceomicans Letzner, 1850; un
~virescens Letzner, 1850; un
~viridicoeruleus Letzner, 1850; un
=psilopterus Kraatz, 1879 "Thüringen & Böhmen"
=?albinae Haury in Géhin, 1885
=niveus Géhin, 1885; nom.pro nivalis Letzner NE-Bohemia:"Riesengebirge"(=Krkonoše Mts.)
~arcticus Sparre-Schneider, 1888, nec Olivier, 1795 C-Sweden:Jämtland:Areskutan
=sollicitans Hartet, 1907 ?England
=britannicus Born, 1908 British Isles:?
=ottonis Csiki, 1909 S-Norway:Trondheim
=lindbergi Burkard, 1921 Finland:Helsingfors(=Helsinki)
~duftschmidi Lapouge, 1922, nec Géhin, 1876 Austria:Linz env.
~pseudocandisatus Eidam, 1927; un
~pseudolaevigatus Breuning, 1934; un Moravia:"Prossnitz"(=Proseènice)
=carelicus Hellén, 1934 Karelia:?Kivenkappa(currently:NW-Russia:W of Ladozhskoe Lake)
=subpurpurascens Eidam, 1941; nom.pro duftschmidi Lapouge Austria:Linz env.
~blumenthali Blumenthal, 1961; un NC-Germany:Lüneburg:Lüneburger Heide:Königsmoor nr.Tostedt
~pseudarcticus Mandl, 1962; un LNF(?N-Finland, Sweden & Norway)
~pseudolindbergi Mandl, 1962; un LNF(?Sweden:Uppland)
~pseudottonis Mandl, 1962; un LNF(?S-Sweden)
~subcarbonatus Niedl, 1975; un S-Bohemia:Mt.Kle nr.Èeský Krumlov
=lindrothi Silfverberg, 1977; nom.pro arcticus Sparre-Schneider C-Sweden:(Jämtland:Areskutan); Kola Peninsula
~intermarinus Blumenthal & Staven, 1985; un NC-Germany:Schleswig-Holstein:Osterwohld W of Albersdorf
=browni Deuve, 1999 NW-Scotland:Raasay Island

ssp. andrzejuscii Fischer, 1823 C,E-Poland(Wolhynien"), NE-Slovakia, CW-Russia, Ukr., Latv., Lith., Est.; NE-Rom., ?Mold.
=carbonatus Schaufuss, 1882 "Podolien"
=subpolitus Eidam, 1924 "Podolien"(=W-Ukraine)
=alterviolaceus Kolbe, 1925 "?Norddeutschland:E of Oder"

ssp. wolffii Dejean, 1826 NW,C,SW-Romania(Banat); E-Hungary; ?SE-Slovakia
=mehelyi Ganglbauer, 1896 NC-Romania:Rodna Mts.
=dacoromanus Hormuzaki, 1903 SW-Romania:Mehedinti
=vlascanus Born, 1909 Walachei(=S-Romania)
~macilentus Petri, 1912, nec Olivier, 1795 "Schässburg, Bistra"(=Sighiºoara, Bistra)
~pygmaeus Petri, 1912; un Romania:?; alpine form
~lucidulus Breuning, 1934; un; nom.pro macilentus Petri "Schässburg, Bistra"(=Sighiºoara, Bistra)
=betuliae Csiki, 1940 NE-Hungary:Szabolcs-Szatmár:Nyírség lowland
~balanensis Csiki, 1942; un CE-Romania:Mts.nr.Balan:Mt.Nagy-Hagymás(=Haghimas Mts. nr.Balan)
~ormayanus Csiki, 1944; un C-Romania:from Cibin Mts.to Szöreny Mts.(=Godeanu, Tarcu)
~porolissensis Csiki, 1944; un NW-Romania:from left Someº bank S of Rodna Mts.to W-slope of Bihor Mts.
~serrorum Csiki, 1944; un C-Romania:Bucegi to Fagaraº Mts.

ssp. salisburgensis Kraatz, 1879 Austria:Salzburg, Oberöst., Vorarlberg; S-Germany; N,C-Switzerland; CE-France:Doubs, Jura
=meyeri Born, 1898 Switzerland:Jura:Creux du Vau
~hermanni Born, 1901, nec hermani Mannerheim, 1827en
=muellerianus Born, 1902 Vorarlberg:Hochfreschen
=pauli Kolbe, 1923 ?Simmentaler Alpen
=sachianus Mandl, 1981 C-Austria:E of Salzburg:Sensengebirge:Feichtau-Alm, 1400m

ssp. purpurascens Fabricius, 1787 C,N-France, Belgium, Holland; Switzerland; NW,C,S-Germany; ?LNF
=coeruleus Voet, 1778; spn
=?curtatus Villers, 1789 ?N-France
=crenatus Sturm, 1815 "Österreich"
=asperulus Kraatz, 1878
=palliardii Gradl, 1881
=suffriani Westhoff, 1881 "Westphalen"
=scaber Schaufuss, 1882 "Hessen"
~subcrenatus Géhin, 1885; un ?Jura
=asperipennis Lapouge, 1921
~laevicostatus Lapouge, 1921, nec Schulz, 1900 ?Belgium:Ft.de Soignes
=crenatocostatus Lapouge, 1922 N-France:?Bas-Rhin
=osismius Lapouge, 1922 CW-France:Cotes-du-Nord
~chatenayi Sirguey, 1931; un
~dayremi Sirguey, 1931; un
~pseudopurpurascens Breuning, 1934; un; nom.pro laevicostatus Lap. ?Belgium:Ft.de Soignes
~uriensis Breuning, 1934; un C-Switzerland:Uri:Reuss valley nr.Erstfeld
~cyaneocinctus Cleu, 1937; un Allier:?
~perraudini Mandl, 1962; un SW-Germany:Baden:Schwarzwald:Nimburg
~gauthieri Cleu, 1968; un Drôme:Foret de Saou(W-part); TNF
~rectigranosus Cleu, 1968; un
~jurassicus (David & Marchal, 1968), nec Born, 1911
=perturbatocostatus (Grotz & Grotz, 1980) Belgium:Liége:Rocherath
~culotianus (Le Mao, 1986); un
~monnayensis (Branger, 1986); un
~enigmaticus (Devecis, 1988); un

ssp. fulgens Charpentier, 1825 Pyrenees; NC,NE-Spain:Burgos to Barcelona
~muelleri Haury, 1878, nec Herbst, 1784 Barcelona env.
~nigrinus Nicolas, 1898; un Hautes-Pyrénées:Pic de Néré nr.Baréges
=pseudofulgens Born, 1905 NC-Spain:?Zaragoza; ?"S of Pyrennees:Logrono, Calahorra"
=gerundensis Born, 1916 NE-Spain:Gerona:Setcasas; SC-France:Pyrenees-Orientales
=pseudomuelleri Codina, 1921 Barcelona, Lerida; alpine form; ?Lerida
=ligneus Sirguey, 1931 ?Lerida
~parallelus Sirguey, 1931; un
=subfulgens Jeanne, 1969 Pyrénées-Atlantiques:Pic d’Orhy, 2000m
~ibericus (Jeanne, 1976), nec Fischer, 1823 "Pyrénées"

ssp. aurichalceus Kraatz, 1879 N-Spain:Santander & Vizcaya(Reinosa & Portugalete nr.Bilbao), Pic.de Europa, Lugo, Oviedo
=asturiensis Born, 1925 Oviedo:Los Cabos
~europae Breuning, 1934; un Picos de Europa:summit area(alpine form)
=lugensis Breuning, 1972 NW-Spain:Lugo:Samos
=marinae Maux, 1995 Palencia:N-part:Castille
~alainsis Meyer, 1996; un Palencia:W of Cervera
~bersoni Maux, 1998; un Palencia:N-part:Castille

ssp. baeterrensis Lapouge, 1901 S-France:Aveyron, Tarn, Aude, Herault(Bessan), Alp.Marit., Gard, Vaucluse, Isere, Ardeche
=provincialis Born, 1903 Alpes-de-Haute-Provence:Digne
=bicinctus Born, 1905 Aude:Ft.de Monthaut, Rennes-les-Bains, Ft.de la Loubatiére
=viridilucens Barthe, 1909 Tarn:Montagnes Noires
~sicardi Sirguey, 1931; un
=thorencicus (Ochs, 1967) Alpes-Maritimes:Loyis du Pin

ssp. exasperatus Duftschmid, 1812 NE,CE-Austria:Graz, Wien env.; S-Moravia; NW,NC-Hungary:Gyor, Budapest env.
=glabrellus Dejean, 1826 "Austria"
=pseudoviolaceus Kraatz, 1886 S-Moravia:"Brünn"(=Brno)
=rakosiensis Csiki, 1905 NC-Hungary:"Rakosfeld"(=Pest:E of Budapest)

ssp. germarii Sturm, 1815 Alps:SC-Switz. - N-Italy - SC,SE-Austria; Slovenia, E-Croatia, Slavonia, S-Hungary; "Krain"
=detritus Drapiez, 1821
=obliquus Thomson, 1875 ?Slavonia:Frushka Gora
=savinicus Hammer, 1906 SC-Austria:Villach env.:"Ober-Seeland"
=rombonensis Bernau, 1911 NW-Slovenia:"Mt.Rombon bei Flitsch"(=Julijske Alpe:Bovec)
~jureceki Sokolar, 1913; un Trentino:Folgaria(=S of Trento)
=ticinensis Born, 1920 "Tessin"(=SC-Switzerland)
=poussielguei Nicolas, 1924 Monti Lessini:Sette Communi(=N of Verona)

ssp. neesi Hoppe & Hornschuch, 1825 SC,E-Austria(Kärnten:Heligenblut) to SW-Switzerland; NE-Italy:Alto Adige:Alpi Atesine
~laevigatus Dejean, 1826, nec Scriba, 1790 Steiermark:Rottenmanner Tauern:alpine zone
=kunzei Heer, 1837
~buendneri Breuning, 1934; un CE-Switzerland:"Graubünden, Bonaduzer Alpen"(=W of Davos:Bonaduz env.)
~styriensis Breuning, 1934; un; nom.pro laevigatus Dejean C-Austria:Steiermark(Rottenmanner Tauern:alpine zone), NE-Kärnten
=pseudoneesi Mandl, 1984 C-Austria:Salzburg:E-part; Steiermark:NW-part(Ramsau, 1300m); alpine form

ssp. vlasuljensis Apfelbeck, 1894 Bosnia(Bosnia/Montenegro:Volujak Mts., Vlasulje; alpine form
=scordiscus Lapouge, 1901 Bosnia
=igmanensis Eidam, 1927 Bosnia:Igman planina
=narentinus Eidam, 1927 Bosnia:E of Narenta:Prenj planina:low altitude
=prenjus Eidam, 1927 Bosnia:Prenj planina:high altitude

ssp. fiorii Born, 1901 NC, NW-Italy(Emilia:Casinalba) to Alpi Liguri; SW-Switz.; E-France:Savoie, Alp.Hte.Prov.
~pedemontanensis Breuning, 1935; un Alpi Liguri:"Viozene"
=dechambrei (Colas, 1952) Alpes-Hte-Provence:Vallé de l’Ubaye:Col de Larche
~adelus Tarrier, 1974; un NW-Italy:Liguria:Imperia:Nava, 800m

ssp. azurescens Dejean, 1826 Croatia(Velebit Mts.:betw.Gospic/Karlobag); Serbia, Bosnia, Montenegro
=krajnensis Born, 1904 CE-Serbia:Krajina, Majdanpek, Negotin
=hercegovinus Born, 1913 Bosnia:Vran Planina, Cvrsnica Planina(=SE of Duvno)
=zabljakensis Eidam, 1927 Montenegro:Durmitor Massif:Zabljak env., Prokletje Mts.
=durmitorus Mandl, 1985 Montenegro:Durmitor Massif; alpine form)
=shljebensis Mandl, 1985 Montenegro/Albania:Prokletje Mts.:Zljeb env.; Mokra gora Mts.

ssp. rilvensis Kolbe, 1887 "Rumelia"; ?SW-Bulgaria, ?Macedonia:?Rila Mts., ?Kozuf Planina(=W of Gevgejija)
~balcanicus Lapouge, 1901, nec Born, 1899 ?W-Balcans
=skombrosensis Eidam, 1927 CW,C-Bulgaria:Rila Mts., Vitosha Massif; C,W-Stara planina Mt.R.
=sofianus Eidam, 1927CW-Bulgaria:Sofia env.

ssp. aurolimbatus Dejean, 1829 C,E-Eur.Russia("Ural"); SW,SC-Siberia; Caucasus, Kazakhstan, ?Kyrgyzstan
=castaneipennis Ménétriés, 1832 "Kaukasus"
=eversmanni Fischer, 1832 "Kirghisensteepe"

ssp. picenus A.& J.B.Villa, 1838 C,S-Italy(Ancona env.)
=romanus Born, 1905 CW-Italy:Lazio:Roma env.(Castelromano)
=authionensis Ochs, 1953 ?Alpes-Maritimes:Authion Massif
=bruschii Tarrier, 1974 C-Italy:Abruzzi:Campo Imperatore, 1800m

ssp. liguriensis Breuning, 1934; ?un NW-Italia:Liguria:Alpi Liguri, Appenine Ligure(Val Bisagno nr.Genova)
=hypochora Tarrier, 1971 Alpes-Maritimes:middle Vésubie valley:Roquebilliere env.

ssp. dryas Gistl, 1857 NE-Greece("Saloniki"=Thesaloniki), SW-Bulgaria, Macedonia, NC-Albania
=merditanus Apfelbeck, 1918 NC-Albania:"Mirdite Mts.:Mt.Maj`e Zebes"(=Mirdite Mts.SW of Bicaj); Gjalica Lumes Mts.
=shardaghensis Apfelbeck, 1918 Macedonia/Kosovo:?Shar Dagh:Golesnica(=Shar Planina)
~skripetarum Breuning, in litt.
=bartoni Maøan, 1930 SW-Bulgaria:"Alibotush"(=Mt.Orvelos)
~korabensis Csiki, 1944; un Macedonia/Serbia/Albania borders:Korab Mts.
~marani Štìrba, 1945; un Macedonia:Jablanica Mts.(=W of Ohrid); Peristeri(=Pelister Mts.SW of Bitola)
~purkynei Štìrba, 1945; un SW-Bulgaria:Belasica Mts.(=nr.Petrich)
~peristericus Mandl, 1961; un Macedonia:"Peristeri"(=Pelister Mts.SW of Bitola)
=letellieri Blumenthal, 1976 NE-Greece:Pangaion Massif, Olympos Massif
=pirinensis Mandl, 1985 SW-Bulgaria:"Pirin:Sandanski-Popinalka"(=Pirin Mts.:S-part:Popina Lyaka:Sandanski Hut)

ssp. mixtus Géhin, 1876, nec Herbst, 1784; nbm CE-France:Isére(Grande Chartreuse); Savoie; Drôme:N-part
=cyaneolimbatus Kraatz, 1879 Isére:Grenoble; "Gallia mer. mont. Costa"(LT)
=saouensis David & Marchal, 1968 Drôme:Ft.de Saou
=neomixtus Tarrier, 1975; nom.pro saouensis David & Marchal Drôme:Ft.de Saou(=E,C-part)
=vocontianus Keith, 1995 Drôme:ca.40km E Valence:Ft.de Lente


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